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Golden Eagle (Aquila chrysaetos) juvenile in flight, Norway, November
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Female Harpy Eagle (Harpia harpyja) in flight. Returning to its nest. Pousada Currupira d Araras, south west Brazil
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Yellow bellied toad (Bombina variegata) showing underside, Kirchheimbolanden, Germany. Meetyourneighbours.net project
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Crested newt (Triturus carnifex) Gelderland, The Netherlands, April. Winner of the Amphibians category of the Wildlife Photographer of the Year Awards (WPOY) Competition 2015
Giant Pacific octopus (Enteroctopus dofleini) experiencing freedom after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Smalltooth sand tiger shark (Odontaspis ferox), view from below. El Hierro. Canary Islands
Short-finned pilot whale (Globicephala macrorhynchus) showing underside whilst vertical underwater, South Tenerife, Canary Islands, Atlantic, October
Indian Cobra or Spectacled Cobra (Naja naja), Karnataka, India
Indian Flying Fox (Pteropus giganteus), flying, India
Yellow Guinea pig standing up and squeaking, against white background
Steppe eagle (Aquila nipalensis) in flight, Oman, November
Barn Owl (Tyto alba) hunting / hovering, Somerset, UK, trained birdBarn Owl (Tyto alba) hunting/hovering, Somerset, UK, trained bird
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Oceanic manta ray (Mobula birostris) accompanied by three Black jacks (Caranx lugubris) silhouetted, Revillagigedo Islands, Mexico, Pacific Ocean
Japanese dwarf flying squirrel (Pteromys volans orii) male sitting in tree, portrait. Hokkaido, Japan. March
Ventral view of Swallowtail butterfly (Papilio machaon), male and female, in flight. The Netherlands. July
Small pearl bordered fritillary butterfly (Boloria selene) resting on flower. The Netherlands. July
Japanese dwarf flying squirrel (Pteromys volans orii) gliding through forest canopy during light snowfall. Hokkaido, Japan. February
Bonnethead shark (Sphyrna tiburo) hunting amongst fish in Turtlegrass (Thalassia testudinum) seagrass bed, view from below
A vibrant Caribbean coral reef with two Reef sharks (Carcharhinus perezi) and Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Galapagos hawk (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Crown daisy (Glebionis coronarium) flowers against sky, low angle view. Cyprus. April
Dwarf lionfish (Dendrochirus brachypterus) opens its awingsa (in fact enlarged pectoralDwarf lionfish (Dendrochirus brachypterus) opens its wings (in fact enlarged pectoral fins) Seraya, Tulamben area, Bali, Indonesia, Java Sea
Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii
Marine iguana (Amblyrhynchus cristatus) swimming back to the surface, Galapagos. Pacific ocean
Gannet (Morus bassanus) adult turning in the air as it begins to plunge dive. Shetland Islands, Scotland, UK, July
Great Frigatebird (Fregata minor) in flight over Midway Atoll. Midway
Great Blue shark (Prionace glauca) viewed from just below the surface, Pico Island, Azores, Portugal, Atlantic Ocean
Roseate Tern (Sterna dougallii) in flight in front of building, Madeira Portugal August
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
RF- Looking up under the gills of toadstools of Porcelain fungus (Oudemansiella mucida), Cornwall, UK
Marsh harrier (Circus aeruginosus) adult female circling over Oilseed rape crop, Norfolk, England, May. Digital composite
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
African fish eagle (Haliaeetus vocifer). Zimanga private game reserve, KwaZulu-Natal, South Africa. June
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
RF - Young lionfish (Pterois volitans) swimming near surface hunting silversides, at dusk. Beacon Rock, Sha ab Mahmood, Sinai, Egypt. Red Sea
Green turtle (Chelonia mydas) from underneath swimming through the surface, Mayotte Island, Comores, Indian Ocean
RF- Bottlenose dolphin (Tursiops truncatus) swimming in a spiral movement, Sandy Ridge, Little Bahama Bank. Bahamas
Hawksbill turtle (Eretmochelys imbricata) ventral view, Philippines, Pacific Ocean. Critically endangered
Mashpi glassfrog (Hyalinobatrachium mashpi) underside, internal organs visible. Captive, occurs in Ecuador
Humpback whale (Megaptera novaeangliae) breaching, Manabi, Ecuador, Pacific Ocean
Three Blacktip reef sharks (Carcharhinus melanopterus) silhouetted just below the ocean surface, Yap, Micronesia, Pacific Ocean
Reef manta ray (Mobula alfredi) swimming with a Twinspot snapper (Lutjanus bohar), Okinawa, Japan, Pacific Ocean