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Young urban Red fox (Vulpes vulpes) standing on a wall at night. Bristol, UK, September. Nominated in the Melvita Nature Images Awards competition 2014
Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
RF - Brown hare (Lepus europaeus) adult in arable field, Scotland, UK, August
Close view of a Green anaconda, (Eunectes murinus) underwater, Formoso River, Bonito, Mato Grosso do Sul, Brazil
Lion (Panthera leo) male standing in cold and rain with strong wind blowing, smelling the air, Masai Mara National Reserve, Kenya
King cobra (Ophiophagus hannah) in strike pose with mouth open, tongue out and glottis (hole like structure in mouth) clearly visible. Malaysia
RF - Raccoon (Procyon lotor) in autumn foliage. Connecticut, USA. October
RF- Male Dog Otter / Eurasian Otter (Lutra lutra) head profile, captive, England
Female Snow leopard (Panthera uncia) with her three month cub looking up at its mother, BioParc Doue la Fontaine, France. Captive
Arctic fox (Alopex lagopus) following scent, in winter pelage. Svalbard, Norway. April
A Perentie (Varanus giganteus) using its forked tongue to detect smells, Alice Springs Reptile Centre, Northern Territory, Australia, Captive. May
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
RF- Orang utan (Pongo pygmaeus) head portrait of dominant male and first orangutan in world to cataracts operated on with eyesight restored
Blue shark (Prionace glauca) close up, Azores, Portugal. Finalist in Wildlife Photographer of the Year Awards (WPOY) Competition 2014
Red Fox (Vulpes vulpes) curled up, with an eye open. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Manta ray (Manta birostris), San Benedicto, Revillagigedo (Socorro) Islands, Mexico, East Pacific Ocean
Lemon sole (Microstomus kitt) portrait. Gardur, south west Iceland. North Atlantic Ocean
Water vole (Arvicola amphibius) standing on hind legs sniffing blackberry, Kent, UK, September
Asiatic elephants (Elephas maximus indicus) sniffing the air as they get close to a water body, Kaziranga National Park, Assam, India
Deer mouse (Peromyscus maniculatus) on giant puffball mushroom, watching mosquito in the moonlight. Blackfoot Valley, Western Montana, USA, July
European Otter (Lutra lutra) walking across snow while sniffing the ground. The Netherlands, December. Captive
Domestic pig, hybrid large white sow in free-range sty, UK, August 2010
Feral domestic rabbit (Oryctolagus cuniculus) Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Bengal Tiger (Panthera tigris tigris) captive, with leaves digitally added
Wild boar (Sus scrofa) piglet, sniffing the air, La Pampa province, Patagonia, Argentina. Captive
Grey seal (Halichoerus grypus) pup and mother sniffing and nuzzling each other. Donna Nook, Lincolnshire, UK. November
Black-headed cat snake (Boiga nigriceps) coiled, portrait, Kubah National Park, Sarawak, Malaysia
Western diamondback rattlesnake (Crotalus atrox) young, coiled up on desert floor at dusk, flicking tongue, Texas, USA. March
European hedgehog (Erinaceus europaeus) walking through forest undergrowth, Finland. September
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Iberian lynx (Lynx pardinus) sniffing scent on dead branch, Parque Natural Sierra de Andujar, Andalucia, Spain. January
RF-Polar bear (Ursus maritimus) swimming in Beaufort Sea, portrait. Near Kaktovik, Arctic National Wildlife Refuge, Alaska, USA. September
RF- Red Admiral butterfly (Vanessa atalanta) perched on Bigtooth Maple (Acer grandidentatum). Lost Maples State Park, Hill Country, Central Texas, USA. November
Giant Anteater (Myrmecophaga tridactyla), two sniffing air in savanna. Caiman Ecological Refuge, Southern Pantanal, Moto Grosso do Sul, Brazil
Scottish Wildcat (Felis sylvestris), captive, UK, June
Spectacled caiman (Caiman crocodilus) resting in shrinking pool, Pouso Alegre lodge. Pantanal, Brazil. Highly commended in the Other Animals Category of the GDT competition 2013
Stewart Island Brown Kiwi / Southern Tokoeka (Apteryx australis lawryi), female probing by scent for sand hoppers in rotting kelp beach wrack, Ocean Beach, Stewart Island, New Zealand
Domestic cat (Felis catus) kitten head portrait. Kokkari harbour, Samos, Eastern Sporades, Greece
Saggy skinned frog (Telmatobius culeus) adult in Lake Titicaca, Andes on the border of Peru and Bolivia
Hippopotamus (Hippopotamus amphibius) ears flicking water when opening after surfacing, Mana Pools National Park, Zimbabwe, October 2012
Two Guillemots (Uria aalge), Vardo, Norway, March
Polar bear (Ursus maritimus) 4-5 year-old on pack ice, low angle shot, off the 1002 area of the Arctic National Wildlife Refuge, North Slope of the Brooks Range, Alaska, Beaufort Sea, autumn
Lion (Panthera leo) male using flehmen response to smell, Masai-Mara Game Reserve, Kenya. Vulnerable species
Close up of eye of Tomato frog {Dyscophus antongili} Maroantsetra, Northeast Madagascar
Bearded pig (Sus barbatus) foraging for food at night, Maliau Basin, Sabah, Borneo
RF- Indian rhinoceros (Rhinoceros unicornis) calf, Kaziranga National Park, Assam, India. Vulnerable species
