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Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
Five white Camargue horses running through the water in Southern France, Europe. May
RF- Quarter horses running in snow at ranch, Shell, Wyoming, USA, February
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
RF- Small group of Pink-footed geese (Anser brachyrhynchus) in flight at dawn, The Wash Estuary, Norfolk, England, UK, October 2011
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Flock of White ibis (Eudocimus albus) in breeding plumage, at rookery on water sFlock of White ibis (Eudocimus albus) in breeding plumage, at rookery on waters edge of the mangrove-covered island, Tampa Bay, Florida, USA, March
Large group of Cavalier King Charles Spaniel, puppies, three of which have tricolour coat and the rest blenheim
Flock of Common Starling (Sturnus vulgaris) gathering before landing at winter roost, Salisbury Plain, Wiltshire, UK, January
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Four Wisent / Bison (Bison bonasus) standing in a row. Bialowieza Forest, Bialowieza National Park, Poland, November
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Big-eye fish (Priacanthus hamrur). Egypt, Red Sea
King penguin (Aptenodytes patagonicus) colony. Right Whale Bay, South Georgia. September
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Sockeye Salmon (Oncorhynchus nerka) travelling upstream on spawning migration, Adams river, British Columbia, Canada, October Taken for the Freshwater Project
School of young European perch (Perca fluviatilis) in Altausseer lake, Austria, July
Masai Giraffe (Giraffa camelopardalis tippelskirchi) juveniles, Masai Mara Game Reserve, Kenya
Turkmenian Wild Ass / Kulan (Equus hemionus kulan) Badkhyz, Turkmenistan. Endangered
Lesser flamingos (Phoeniconaias minor) flock in Lake Bogoria, Rift valley, Kenya
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
Common starling (Sturnus vulgaris) flock at dusk engaging in pre-roost display, Dumfries-shire, Scotland, March
Schooling Scalloped hammerhead sharks (Sphyrna lewini) Cocos Island National Park, Costa Rica
Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December
Atlantic mobula ray (Mobula hypostoma) group swimming in Caribbean Sea. Isla Mujeres, Quintana Roo, Mexico
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Gannet (Morus bassanus) chick in a line of adults. Shetland Islands, Scotland, UK, August
Mute swan (Cygnus olor) flock on snow, Hazerswoude, The Netherlands, February
Oriental sweetlips (Plectorhinchus vittatus), Maldives, Indian Ocean
Demoiselle cranes (Anthropoides virgo) flock, India
Curimbata (Prochilodus lineatus) in a clear stream of the Pantanal, with Water hyacinth (Eichhornia sp.), Mato Grosso do Sul, Brazil. Taken for the Freshwater Project
Grey herons (Ardea cinerea) standing in water a dawn, Puszaszer, Hungary, July
Two African lion (Panthera leo) cubs walking on a path. Okavango Delta, Botswana
Domestic dog, St. Bernard / Alpine Mastiff, three on snow in Alps, France
A group of Pure Arab, Shagya Arab and East Bulgarian fillies running in snow, Kabiuk National Stud, Shumen, Bulgaria
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Lion (Panthera leo) cubs sitting on termite mound, Masai-Mara, Kenya. Vulnerable species
Sooty Terns (Onychoprion fuscata) flock in flight, Midway Island. Central Pacific
Lesser flamingos (Phopeniconaias minor) flock, aerial view, Natron lake, Rift valley, Tanzania, Africa
Lion (Panthera leo) males resting, Masai-Mara Game Reserve, Kenya. Vulnerable species
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Heermanns Gulls (Larus heermanni) in flight. Ensenada, Baja California Peninsula, Mexico, December
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January