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Winter gale in sea at sunset, Long Island Sound, Hammonasset State Park, Madison, Connecticut, USA
Purple gallinule (Porphyrio martinicus) reaches for Giant bulrush seeds (Schoenoplectis californicus). Wakodahatchee Wetlands, Florida, USA, April
Canada goose (Branta canadensis) in flight in autumn, Maryland, USA. October
Queen butterfly (Danaus gilippus) emerging from chrysalis on Aquatic milkweed (Asclepias perennis). Hill Country, Texas, USA. Sequence 1/12
Pine warbler (Dendroica pinus) male perched in blossoming Peach (Prunus persica) tree. Hill Country, Texas, USA
Gulf fritillary butterfly (Agraulis vanillae) expanding wings after emerging from chrysalis. Hill Country, Texas, USA
Blackpoll warbler (Setophaga striata) male perched on Mexican olive (Cordia boissieri). South Padre Island, Texas, USA
Northern harrier (Circus hudsonius), two juvenile females hunting. North Park, Colorado, USA. August
Grand Prismatic Springs on cold winter day with mist / vapour, Yellowstone National Park, USA, January
Leopard houndshark (Triakis semifasciata) group swimming amongst rays of light. San Diego, California, USA
Prairie and eroded formaions at Sage Creek Basin, Badlands National Park, South Dakota, USA. August
Eroded badlands and prairie grasses, Badlands National Park, South Dakota, USA. August
Devils Tower, seen from Joyner Ridge trailhead; Devils Tower National Monument, Wyoming, USA. August
Lemon shark (Negaprion brevirostris) hunting over Turtlegrass (Thalassia testudinum) seagrass bed, view from below. Florida Keys, Florida, USA
Orange garibaldi damselfish (Hypsypops rubicundus) in a giant kelp (Macrocystis pyrifera) forest. Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America
Hickman Bridge, rock arch in Navajo sandstone. Capitol Reef National Park, Utah, USA. February 2020
Least Chipmunk (Tamias minimus) feeding on seeds, Grand Teton National Park, Wyoming, USA
Forest with lush vegetation Portland, Oregon, USA. April
Portrait of a Red-breasted nuthatch, (Sitta canadensis) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Chipping sparrow (Spizella passerina), adult perched in flowering eastern redbud in spring, New York, USA, May
Blue jay (Cyanocitta cristata) in flight carrying an acorn that it will cache for winter food supply, New York, USA, October
Osprey (Pandion haliaetus) fishing. Acadia National Park, Maine, USA. May
North American river otter (Lontra canadensis) on snow covered bank, reflected in icy water. Yellowstone National Park, USA, January
RF - North American river otters (Lontra canadensis) (probably female with two juveniles) on the frozen river edge.Yellowstone National Park, USA. January. Digitally stitched image
RF - Red fox (Vulpes vulpes) on the edge of coniferous forest. Yellowstone National Park, USA. January (This image may be licensed either as rights managed or royalty free.)
RF - Lodgepole pines (Pinus contorta) covered in heavy early morning hoar frost. Yellowstone National Park, USA. January. Digitally stitched image
Red fox (Vulpes vulpes) walking through deep winter snow. Hayden Valley, Yellowstone, USA. January
North American river otter (Lontra canadensis) female and cub walking across snow, reflected in water. Yellowstone National Park, USA, January
Sea otter (Enhydra lutris) female and pup. Alaska, USA, February
Coyote (Canis latrans) standing in snow, dwarfed by frosty trees. Yellowstone National Park, USA, January
Black-tailed rattlesnake (Crotalus molossus), Chiricahua Mountains, Arizona, USA. August
Alpaca (Lama pacos) portrait, New York, USA
Pacific gopher snake (Pituophis melanoleucus catenifer), Captive, USA
Hanging thief robber fly (Diogmites sp) Philadelphia, Pennsylvania, USA, July
Steller sea lions (Eumetopias jubatus) playing underwater, Inian Island, Alaska, USA. August
American red squirrel (Tamiasciurus hudsonicus) on tree trunk drinking in a beaver pond early in the morning. Acadia National Park, Maine, USA. May
Ridgeways rail (Rallus obsoletus levipes) foraging in wetlands, Bolsa Chica Ecological Reserve, California, USA April
California least terns (Sternula antillarum browni) courtship behaviour, Bolsa Chica Ecological Reserve, California, USA June
Blue jays (Cyanocitta cristata) confrontation in snow, April 2020. New York State, USA
Northern Cardinals (Cardinalis cardinalis) male and female in the snow, photographed from low angle, Freeville, New York, USA. April 2020
Yellow Warbler (Setophaga petechia, formerly Dendroica petechia), male in breeding plumage, Ithaca, New York, USA. May 2020
Canada goose (Branta canadensis) with gosling, a few days old. Massachusetts, USA. April
Wood duck (Aix sponsa) male in breeding plumage. Acadia National Park, Maine, USA. April
Long-beaked common dolphin (Delphinus capensis) off the coast of California, United States
Common Merganser (Mergus merganser) female with ducklings, in threat posture at the approach of a second female, Lansing, New York, USA
Baltimore Oriole (Icterus galbula) male with spider prey, perched in crabapple blossom in spring, Ithaca, New York, USA
Purple Finch (Carpodacus purpureus) perched amid apple blossom, New York, USA