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Blue-footed booby (Sula nebouxii) courting pair, South coast, Santa Cruz Island, Galapagos
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Blue-footed booby (Sula nebouxii) landing, South coast, Santa Cruz Island, Galapagos
Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Giants Causeway, columnar basalt columns, result of an ancient volcanic fissure eruption, Northern Ireland, UK, World Heritage site. September 2019
Red-footed booby (Sula sula) in flight with wings outstretched. Genovesa Island, Galapagos
Underwater view of pebbles near the shore of Lake Baikal Lake Baikal UNESCO World Heritage Site, Siberia, Russia. May 2013. Photographed for The Freshwater Project
Green tree ants (Oecophylla smaragdina) defending their leaf nest, Daintree River, Wet Tropics World Heritage area, Queensland, Australia
White-lipped green tree frog (Litoria infrafrenata) clinging on to a branch, Etty Bay, Wet Tropics World Heritage area, Queensland, Australia
Aerial view of the front of the Sermeq Kujalleq Glacier, Greenland, entering the Kangia Ilulissat Icefjord full of icebergs
Short-beaked echidna (Tachyglossus aculeatus). Francois Peron National Park, Shark Bay, Western Australia. October
Marine iguanas (Amblyrhynchus cristatus) silhouettes at sunset, Punta Vicente Roca, Isabela Island, Galapagos
Aerial view of the Joggins Fossil Cliffs UNESCO World Heritage Site along the shore of the Bay of Fundy, Nova Scotia, Canada. May 2017
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Green turtle (Chelonia mydas) swimming near mangroves, Elizabeth Bay, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Rice (Oryza sativa) terraces. Jatiluwih Green Land, Bali, Indonesia. 2015
Palms growing in front of Rice (Oryza sativa) terrace. Jatiluwih Green Land, Bali, Indonesia. 2015
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
The Lagoon Coral Reef, Lord Howe IslandThis breathtaking photo print captures the essence of this UNESCO Natural World Heritage Site in all its glory
Galapagos sea lion (Zalophus wollebaeki) yearling pup playing withg own air bubbles in shallows, Gardner Islet, Espanola Island, Galapagos, Ecuador
Gaur (Bos gaurus) Manas National Park UNESCO World Heritage Site, Assam, India
Two Red-footed booby (Sula sula) perched on branches, Genovesa Island, Galapagos Islands, Ecuador
Close up of a silverback Mountain gorilla (Gorilla beringei beringei) face with eyes closed, and hand / fingers, Virunga National Park, Democratic Republic of Congo
Male Blue-footed booby (Sula nebouxii) walking on sand, San Cristobal Island, Galapagos, South America
Galapagos penguin (Spheniscus mendiculus), pair courting, Isabela Island, Galapagos Islands
Spotted eagle ray (Aetobatus narinari), Kicker Rock, San Cristobal Island, Galapagos
Sandstone rock formations in the Sahara desert. Ennedi Natural and Cultural Reserve, UNESCO World Heritage Site, Chad. September 2019
Vampire ground finch (Geospiza septentrionalis) attacking large chick to drink blood, after it was wounded by aggressive adult booby, Wolf Island, Galapagos
Pinzon giant tortoise (Chelonoidis duncanensis), saddleback type typical of arid island, their long necks and raised shell allowing them to browse on cacti
Spotted eagle ray (Aetobatus narinari) Kicker Rock, San Cristobal Island, Galapagos
Galapagos penguins (Spheniscus mendiculus) underwater with shoal of fish, Tagus Cove, Isabela Island, Galapagos
Galapagos hawk (Buteo galapagoensis) flying, carrying an iguana carcass, Punta Espinosa, Fernandina Island, Galapagos
Floreana giant tortoise hybrid descendant (Chelonoidis elephantopus), Fausto Llerena Giant Trotoise Breeding Centre, Galapagos National park Directorate, Puerto Ayora, Santa Cruz Island, Galapagos
Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Sally lightfoot crabs (Grapsus grapsus) on rock with waves, Cape Douglas, Fernandina Island, Galapagos
Tibetan gazelle (Procapra picticaudata) in snow. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Army ant (Eciton burchellii) major soldier, Parque da Onca Parda Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Transparent black ice with cracks on Lake Baikal, aerial shot. Photographed for The Freshwater Project extended. Near Khuzhir, Olchon Island, Irkutsk Oblast and Buryat Republic, Siberia, Russia
Red-footed booby (Sula sula), adult and chick at nest. Genovesa Island, Galapagos
Nazca booby (Sula granti) preening. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Marine iguana (Amblyrhynchus cristatus), Punta Vicente Roca, Isabela Island, Galapagos
Green turtle (Chelonia mydas) female laying eggs, North coast, Santa Cruz Island, Galapagos
Northern rockhopper penguin (Eudyptes moseleyi) parent with chick. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Marine iguana (Amblyrhynchus cristatus) basking on volcanic rock at coast. Isabela Island, Galapagos
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti) at nest with egg. Overlooking sea, Gardner Islet, Floreana Island, Galapagos. December 2014
Nazca booby (Sula granti), pair amongst Prickly pear (Opuntia sp) cacti at coast. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Flowers (Trollius sp) Balang Mountain, Wolong National Nature Reserve, Sichuan Giant Panda Sanctuaries - Wolong, Mt Siguniang and Jiajin Mountains UNESCO World Heritage Site, Sichuan Province, China
Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo
Norfolk Island pine trees (Araucaria heterophylla) near lagoon with Mount Lidgbird and Mount Gower in the background, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island
Low angle view of Porcelain fungus (Oudemansiella mucida) growing on a dead Beech tree (Fagus sylvatica). Plitvice Lakes National Park, Croatia. November
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Thomass Langur (Presbytis thomasi), animal endemic to northern Sumatra. These primates have relatively long arms and legs, adapted for an acrobatic life in the crowns of trees
Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water to keep cool, Alcedo Volcano, Isabela Island, Galapagos
Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) males in breeding colouration on rocks, with sun halo, Floreana Island, Galapagos
Green turtle (Chelonia mydas), female returning to the sea after egg laying, Puerto Pajas, Isabela Island, Galapagos
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Remoras (Remora sp. ) hitchhiking on the tail of a large (13-15m) female whale sharkRemoras (Remora sp.) hitchhiking on the tail of a large (13-15m) female whale shark (Rhincodon typus) swims in open water. Wolf Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
River running through cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Skelleg Michael Monastery Steps, County Kerry, Republic of Ireland. June 2014
Portland Bill Lighthouse, Isle of Portland, Dorset, England, UK. November 2011
Mountain gorilla (Gorilla beringei beringei) female resting with her baby, members of the Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Mountain gorillas (Gorilla gorilla beringei) grooming, members of the Munyaga group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered