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Gravid female Glass / Ghost shrimp on Glass sponge (Hexactinellida) from coral seamount, SW Indian Ridge, Indian Ocean, December 2011
Water fleas (Daphnia sp. ) and a green algae (Volvox aureus) in water from a garden pondWater fleas (Daphnia sp.) and a green algae (Volvox aureus) in water from a garden pond. Derbyshire, UK. September. Digital composite
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Dolphin trainer interacting with Bottlenose Dolphin (Tursiops truncatus), Dolphin Reef, Eilat, Israel - Red Sea. Model released
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Puffin (Fratercula arctica) swimming underwater, Farne Islands, Northumberland, UK, July
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Roughtail stingray (Dasyatis centura) on sea floor, Tenerife, Canary Islands
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
Humpback whale (Megaptera novaeangliae) male calf relaxing with his mother and an escort whale in the background. Vava'u, Tonga, South Pacific
Ripples of sand on a shallow seabed. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Humpback whale (Megaptera novaeangliae) just under surface, off Shetland, Scotland, UK. First ever underwater images of humpbacks in British waters
Kelp forest (Laminaria digitata) with small fish, Shetland, Scotland, UK, July
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Marine iguana (Amblyrhynchus cristatus) swimming underwater, Fernandina Island, Galapagos
Fin whale (Balaenoptera physalus) near surface. Pico Island, Azores, Portugal. May
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Florida manatee (Trichechus manatus latirostris) swimming into a fresh water spring, Three Sisters Spring, Crystal River, Florida, USA
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Underwater view of Waterlilies (Nymphaea alba) in a lake. Alps, Ain, France, June
Rainbow river, Rainbow Springs State Park, Florida, USA. January. Photographed for The Freshwater Project
Sunrays in Cenote Jardin del Eden, Quintana Roo, Yucatan Peninsula, Mexico, May 2016. Photographed for The Freshwater Project
Yellow bellied toad (Bombina variegata) showing underside, Kirchheimbolanden, Germany. Meetyourneighbours.net project
Caribbean reef squid (Sepioteuthis sepioidea) in mid water over a coral reef. East End, Grand Cayman, Cayman Islands, West Indies. Caribbean Sea
Spotted Wobbegong Shark (Orectolobus maculatus) lying in seaweed. Manly, Sydney, New South Wales, Australia. Pacific Ocean. November
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA
Atlantic halibut (Hippoglossus hippoglossus) portrait, Saltstraumen, Bod, Norway, October 2008
Nudibranch (Flabellina nobilis) photographed in the field aquarium in Gulen, Norway. North East Atlantic Ocean
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
A Florida manatee (Trichechus manatus latirostrus) head portrait in Three Sisters Spring. Crystal River, Florida, USA. February 2010
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Underwater view of pebbles near the shore of Lake Baikal Lake Baikal UNESCO World Heritage Site, Siberia, Russia. May 2013. Photographed for The Freshwater Project
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands
Green turtle (Chelonia mydas) feeding on on Turtlegrass (Thalassia testudinum) seagrass bed. The Bahamas
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Common toad (Bufo bufo) tadpole swimming seen from below, Belgium, June
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Brown bear (Ursus arctos) paw seem from under water, Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia, August
Green turtle (Chelonia mydas) with rays of sunlight, black and white image, Akumal, Caribbean Sea, Mexico, July. Second place in the Visions of our Nature 2018
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June. Photographed for The Freshwater Project
Illuminated netdevil (Linophyne arboritera), male anglerfish attached to much larger female, captive from deep sea
Bream (Abramis brama) and European chub (Leuciscus / Squalius cephalus) shoal in a calm part of the Rhine, Schaffhausen, Switzerland, February 2009 WWE OUTDOOR EXHIBITION