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Fingals Cave, showing basalt columns, Isle of Staffa, Inner Hebrides, Scotland, UK, June 2010
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Split level landscape in oxbow of the Aare river, Switzerland, September
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Ice formations, Lake Sassolo, Sambuco valley, Ticino, Switzerland
Plumose sea anemones (Metridium senile) on a ship wreck, Lofoten, Norway, November 2008
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Waves lashing the Four Lighthouse during winter storm, Northern Brittany, France, December 2011. Sequence 8/12
Lilac-breasted roller (Coracias caudata) stretching wings on acacia branch, Masai Mara Game Reserve, Kenya. November
Southern stingray (Hypanus americanus) female swimming over a shallow sand bank, under morning sun, North Sound, Grand Cayman, Cayman Islands. British West Indies, Caribbean Sea
Palm tree branch dipping towards water, Puerto Jimenez, Golfo Dulce, Osa Peninsula, Costa Rica
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Underwater scene where light and water interplay on a shallow sand bank, Grand Cayman, Cayman Islands, British West Indies, Caribbean Sea, September 2005
Young Florida manatee (Trichechus manatus latirostrus) portrait, Three Sisters Spring, Crystal River, Florida, USA. February 2010
Southern stingray (Hypanus americanus) swimming over sand ripples on sandbar, Stingray City, Grand Cayman. British West Indies
Free diver coming up from dive to explore coral reef, Sinai, Egypt, Red Sea Model released
Lago di Landro at dawn, Dolomite Mountains, Sud Tirol / Alto Adige, ItalyLago di Landro at dawn, Dolomite Mountains, Sud Tirol/Alto Adige, Italy
Large pod of Bottlenose dolphins (Tursiops truncatus) porpoising over waves during annual sardine run, Port St Johns, South Africa
Two Atlantic spotted dolphins (Stenella frontalis) over a shallow sand bank. On the left is a juvenile which does not have the characteristic spots of the adults
Crystal clear, shallow seas, Porto Pino, Sardinia, Mediterranean Sea
Sand ripples on seabed and sun burst in shallow, clear water. Capo Comino, Sardinia, Mediterranean Sea
Cairns birdwing butterfly (Ornithoptera euphorion) male resting on fern, large butterfly endemic to wet tropics in far north Queensland, Cairns, Queensland, Australia
Green sea turtle (Chelonia mydas) near surface in shallow water. The Bahamas
Aerial view of Southern right whale (Eubalaena australis) female with calf in shallow coastal water, South Africa, Indian Ocean
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Laguna 69, Huascaran National Park, Andes, Peru, November 2013
Detail of small waves on the eastern Atlantic. Isle of Coll, Scotland, October 2012
River Lepenjica flowing through narrow gap in rocks, Triglav National Park, Slovenia, June 2009
River Lepenjica, with a pothole in rock, Triglav National Park, Slovenia, June 2009
Forest kelp (Laminaria hyperborea) Saltstraumen, Bod, Norway, October 2008
Galapagos sea lion (Zalophus wollebaeki) emerging from water, Galapagos
Green waters of Emerald Lake, near Carcross, Yukon Territories, Canada, September 2013
Detail of the pectoral fin of Roundhead parrotfish (Scarus viridifucatus) coast of Dhofar and Hallaniyat islands, Oman. Arabian Sea
Bowhead Whale (Balaena mysticetus) rubbing off flaking skin on the ocean bottom, east coast Baffin island, Qikiqtarjuaq, Nunavut, Canada, May
American Purple Gallinule (Porphyrio martinica) displaying brightly coloured wings. Everglades National Park, Florida, USA, March
Scenic view of a bull kelp forest (Nereocystis luetkeana) with sunlight shining through the fronds. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Bigeye trevally or jack (Caranx sexfasciatus) shoal, Sipadan, Malaysia
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Southern right whale (Eubalaena australis) with calluses covered in parasitic crustaceans named cyamids or whale lice (Cyamus ovalis)
Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs
Isolated rain storm above the sea. Bahamas Sea, Bahamas, North America, August 2006
A young Grey Seal (Halichoerus grypus) above kelp in the Farne Islands. This seal has an orange snout from foraging on a rusty ship wreck. Northumberland, England, UK. North Sea. February
Long exposure image of Tiger shark (Galeocerdo cuvier) chasing bait, Bahamas
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
Underwater landscape in Aare river, Canton of Berne, Switzerland, January
Baby Olive Ridley sea turtle (Lepidochelys olivacea) struggles against the swell to swim away from the beach where it hatched, Ostional, Costa Rica. October