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Portrait of a Blackbar soldierfish (Myripristis jacobus) infected with Cymothoid isopod parasites. The large parasite between the eyes is a female, the smaller individuals are males
Pair of Rock beauty angelfish (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of the female during the spawning rise
Veined octopus (Amphioctopus marginatus) sheltering in an old clam shell on the sandy seabed. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
View through a cavern in a coral reef, with a Small grouper (Cephalopholis cruentata) in the distance. Snapper Hole, East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
View of steering wheel inside Bedford OYC water tanker 3 ton lorry, from another Bedford OYC truck, deep inside hold of wreck of HMS Thistlegorm
Spotted eagle rays (Aetobatus narinari), in mangrove inlet, Elizabeth Bay, Isabela Island, Galapagos
Marbled Stingray (Taeniurops meyeni), Galapagos
Lemon shark pups (Negaprion brevirostris) in mangrove forest, Eleuthera, Bahamas
Toucan barbet (Semnornis ramphastinus) Bellavista cloud forest private reserve, 1700m altitude, Tandayapa Valley, Andean cloud forest, Ecuador
Humpback whale (Megaptera novaeangliae) breaching, Kwazulu-Natal, South Africa
Split level with Table corals (Acropora spp. ) growing close to an islandSplit level with Table corals (Acropora spp.) growing close to an island. Pelee Island, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Diverse hard coral growth (Pocillopora damicornis, Pocillopora verrucosa and Millepora dichotoma), flourishes beneath the barren desert cliffs in the Red Sea. Ras Katy, Sinai, Egypt
Stingless golden jellyfish (Mastigias sp. ), backlit by the sunStingless golden jellyfish (Mastigias sp.), backlit by the sun, in a landlocked marine lake in the middle of an island. Their golden colour comes from endosymbiotic algae
Group of Longfin batfish (Platax teira) beneath the surface, close to an island. Farondi Islands, Misool, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Tiny snapping shrimp (Synalpheus sp. ) looks out from within sea squirt (Didemnum molleTiny snapping shrimp (Synalpheus sp.) looks out from within sea squirt (Didemnum molle) at night on a coral reef. Piaynemo Island, Raja Ampat, West Papua, Indonesia
Saddleback anemonefish (Amphiprion polymnus) barks a warning as it guards a clutch of developing eggs (beneath it) laid on a small stone, below a fold in its home
Sunbeams play through the roots of red mangrove trees (Rhizophora sp.), encrusted with marine life and home to schoolmaster snappers (Lutjanus apodus)
Caribbean reef shark (Carcharhinus perezi) swims over a coral reef, photographed through the branches of a porous sea rod (Pseudoplexaura sp.)
A group of large stingrays (Dasyatis americana) swim over sand in shallow water, in a split level photo taken at dawn. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Yellowhead jawfish (Opistognathus aurifrons) male rises up from its burrow and aerates the eggs he is mouthbrooding. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Orange branching sponge (Pilocaulis sp. ) growing in a coral canyon on a reef wallOrange branching sponge (Pilocaulis sp.) growing in a coral canyon on a reef wall. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Slender filefish (Monacanthus tuckeri) hiding in the branches of a porous sea rod (Pseudoplexaura sp.) on a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - American crocodile (Crocodylus acutus) swimming through sunrays, above a bed of seagrass ( Thalassia testudinum). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - A threespot damselfish (Stegastes planifrons) aggressively defending its algae garden on a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Model Released
RF - Silhouette of a large Pelagic silky shark (Carcharhinus falciformis) beneath a sunburst. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Fin whale (Balaenoptera physalus), two feeding, aerial view. Baja California, Mexico. March
Grey whale (Eschrichtius robustus) pod migrating north, aerial view. Baja California, Mexico. February
Humpback whale (Megaptera novaeangliae) spouting, rainbow effect created in spray, aerial view. Baja California, Mexico. March
RF - Banggai cardinalfish (Pterapogon kauderni). Lembeh Strait, North Sulawesi, Indonesia. (This image may be licensed either as rights managed or royalty free.)
Zebra / leopard shark (Stegostoma fasciatum) swimming over the reef, Vaavu atoll, Maldives, Indian Ocean
Northern muriqui monkey (Brachyteles hypoxanthus) hanging in a tree, RPPN Feliciano Miguel Abdala, Atlantic Forest, Brazil. June
Long-wattled umbrellabird (Cephalopterus penduliger) Buenaventura Reserve, Ecuador
Coral heads surrounded by Blacktip cardinalfish (Apogon atradorsatus)and juvenile salemas (Xenocys jessiae), Rabida Island, Galapagos
Blacktip shark (Carcharhinus limbatus) pup in a tranquil inlet, Santa Cruz. Tortuga Bay, Santa Cruz Island, Galapagos
Green turtle (Chelonia mydas) swimming with shoal of striped salemas, Kicker Rock, San Cristobal Island, Galapagos
Golden Rays (Rhinoptera steindachneri) swimming with Green turtle (Chenolia mydas). Isabela Island. Galapagos
Galapagos sea lion (Zalophus wollebaeki) hunting, Champion Islet, Floreana Island, Galapagos
Bonito (Sarda orientalis) shoal swimming past the undersea base of Darwinas ArchBonito (Sarda orientalis) shoal swimming past the undersea base of Darwins Arch. Darwin (Culpepper) Island, Galapagos
Green turtle (Chelonia mydas) swimming, Posa de los Patillos, Fernandina Island, Galapagos
Galapagos sea lions (Zalophus wollebaeki) sub adult males playing with starfish, Mosquera Islet, Galapagos
Flightless cormorant (Phalacrocorax harrisi) coming ashore amongst crabs. Clearwater Bay, Isabela Island, Galapagos. Sequence 3 of 3
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Bicolor damselfish (Stegastes partitus), Guanahacabibes Peninsula National Park, Pinar del Rio Province, western Cuba
Foureye butterflyfish (Chaetodon capistratus) couple feeding on coral, Guanahacabibes Peninsula National Park, Pinar del Rio Province, western Cuba
Schoolmaster snapper (Lutjanus apodus) and Bluestriped grunt (Haemulon sciurus) seeking protection in Elkhorn Coral (Acropora palmata), Jardines de la Reina / Gardens of the Queen National Park
Silky Shark (Carcharhinus falciformis) with hook, Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba
Black grouper (Mycteroperca bonaci), and Caribbean Reef Shark (Carcharhinus perezi), Jardines de la Reina / Gardens of the Queen National Park, Caribbean Sea, Ciego de Avila, Cuba