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Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Hawksbill turtle (Eretmochelys imbricata) swimming underwater, Nosy Be, North Madagascar
Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Aerial view of boats and snorkelers, Looe Key Reef, Florida Keys National Marine Sanctuary, USA, showing spur and groove coral formations
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
American crocodile (Crocodylus acutus) shows off its teeth. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Close up portrait of the face of a Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
Inflated spotted porcupinefish (Diodon hystrix), Maui, Hawaii
Pod of Indian Ocean bottlenose dolphin (Tursiops adunctus) swim over a coral reef. Sha ab El Erg, Hurghada, Egypt. Red Sea
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo. Photographed with natural light and UV light
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Gravid female Glass / Ghost shrimp on Glass sponge (Hexactinellida) from coral seamount, SW Indian Ridge, Indian Ocean, December 2011
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
Humpback whale (Megaptera novaeangliae) male calf relaxing with his mother and an escort whale in the background. Vava'u, Tonga, South Pacific
Ripples of sand on a shallow seabed. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Caribbean reef squid (Sepioteuthis sepioidea) in mid water over a coral reef. East End, Grand Cayman, Cayman Islands, West Indies. Caribbean Sea
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Brown throated Three-toed Sloth (Bradypus variegatus) mother and newborn baby (less than 1 week old) Aviarios Sloth Sanctuary, Costa Rica *Digitally removed highlights in background
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Green turtle (Chelonia mydas) feeding on on Turtlegrass (Thalassia testudinum) seagrass bed. The Bahamas
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Green turtle (Chelonia mydas) with rays of sunlight, black and white image, Akumal, Caribbean Sea, Mexico, July. Second place in the Visions of our Nature 2018
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula