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RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011
Hawksbill turtle (Eretmochelys imbricata) swimming underwater, Nosy Be, North Madagascar
Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Underwater river scene with freshwater plants and Tetra fish, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Hairy frogfish / Striated frogfish (Antennarius striatus), portrait, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Inflated spotted porcupinefish (Diodon hystrix), Maui, Hawaii
Pod of Indian Ocean bottlenose dolphin (Tursiops adunctus) swim over a coral reef. Sha ab El Erg, Hurghada, Egypt. Red Sea
American crocodile (Crocodylus acutus) shows off its teeth. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Great hammerhead shark (Sphyrna mokarran) swimming over a sandy seabed, Bimini, Bahamas
Close up portrait of the face of a Nurse shark (Ginglymostoma cirratum) resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico
Pair of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
RF - American crocodile (Crocodylus acutus) swimming through sunrays, above a bed of seagrass ( Thalassia testudinum). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
A shiver of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef with Common sea fans (Gorgonia ventalina) and sea plumes (Pseudopterogorgia sp)
Borneo giant forest scorpion (Heterometrus longimanus) resting inside a fallen hollow log. Danum Valley, Sabah, Borneo
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Gravid female Glass / Ghost shrimp on Glass sponge (Hexactinellida) from coral seamount, SW Indian Ridge, Indian Ocean, December 2011
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Great white shark (Carcharodon carcharias) underwater, Guadalupe Island, Mexico (North Pacific)
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Electric fileclam / Disco clam (Ctenoides ales). West Papua, Indonesia. Indo-West Pacific
Colorful Pfeffers flamboyant cuttlefish (Metasepia pfefferi) swimming over a colony of cup coral at night, Philippines
Ripples of sand on a shallow seabed. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Green turtle (Chelonia mydas) with rays of sunlight, black and white image, Akumal, Caribbean Sea, Mexico, July. Second place in the Visions of our Nature 2018
Caribbean reef squid (Sepioteuthis sepioidea) in mid water over a coral reef. East End, Grand Cayman, Cayman Islands, West Indies. Caribbean Sea
Florida manatee (Trichechus manatus latirostris) at Three Sisters Spring in Crystal River, Florida, USA
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
RF- Western clownfish (Amphiprion oceallaris) in Magnificent sea anemone (Heteractis magnifica). Raja Ampat, West Papua, Indonesia
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Brown throated Three-toed Sloth (Bradypus variegatus) mother and newborn baby (less than 1 week old) Aviarios Sloth Sanctuary, Costa Rica *Digitally removed highlights in background
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Bulb tentacle sea anemone (Entacmaea quadricolor) close up detail, Maldives, Indian Ocean
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Green turtle (Chelonia mydas) feeding on on Turtlegrass (Thalassia testudinum) seagrass bed. The Bahamas
Green turtle (Chelonia mydas) swimming near mangroves, Elizabeth Bay, Isabela Island, Galapagos
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
A wide angle macro view of a Veined octopus ( Amphioctopus marginatus) stretching out from its protective den in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Three Sperm whales (Physeter macrocephalus) socialising under the surface, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species
Aerial view of boats and snorkelers, Looe Key Reef, Florida Keys National Marine Sanctuary, USA, showing spur and groove coral formations
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp)Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Healthy coral reef landscape, Carl's Ultimate dive site in the Eastern Fields of Papua New Guinea
Chestnut crowned antpitta (Grallaria ruficapillas), Mindo, Pichincha, Ecuador
Galapagos sea lion (Zalophus wollebaeki) yearling pup playing withg own air bubbles in shallows, Gardner Islet, Espanola Island, Galapagos, Ecuador
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
California sea lions (Zalophus californianus) playing in a kelp forest off Santa Barbara Island, California, USA
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
RF Humpback whale (Megaptera novaeangliae) aerial view. Baja California, Mexico
Pygmy blue whale (Balaenoptera musculus brevicauda) subspecies of blue whale, diving downwards, Mirissa, Sri Lanka, Indian Ocean. Endangered species
A sprouting coconut floating in the sea close to the shore, Yap, Micronesia, Pacific Ocean
Tail-spot blenny (Ecsenius stigmatura) peering out from amongst the coral, Triton Bay, West Papua, Indonesia, Pacific Ocean
Female Peacock-mantis shrimp (Odontodactylus scyllarus) carrying her eggs, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Frill-necked lizard (Chlamydosaurus kingii) threat display with neck frill extended, Cooktown, Queensland, Australia