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Red deer (Cervus elaphus) hind in frost covered pine forest, Cairngorms National Park, Highlands, Scotland, UK, October
Red squirrel (Sciurus vulgaris) sitting on an old giant gnarled Oak tree. Highlands, Scotland, UK, January
Red squirrels (Sciurus vulgaris) three chasing each other round oak tree, Cairngorms National Park, Highlands, Scotland, UK, April
Scots Pine trees (Pinus sylvestris) reflected in loch, Abernethy Forest, Scotland, UK, November
Scots pines (Pinus sylvestris) and regeneration on moorland, Abernethy, Cairngorms National Park, Scotland, UK, November 2011
Elevated view over lochs and pine forest, Rothiemurchus, Cairngorms National Park, Scotland, UK, September 2013
Loch Garten and surrounding pine forest in morning light, Abernethy, Cairngorms National Park, Scotland, UK, September 2013
Red deer (Cervus elaphus) hinds leaving shelter of forest in winter, Scotland, UK, February
Scots pine (Pinus sylvestris) trees growing in wooded ravine, Beinn Eighe National Nature Reserve, Wester Ross, Scotland, UK, October 2015
Silver birch (Betula pendula) woodland in early autumn, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, October 2014
Herd of Red deer (Cervus elaphus) stags in clearing in snow covered pine forest, Cairngorms National Park, Scotland, UK, December
Scots pine (Pinus sylvestris) sapling on heather moorland, Strathspey, Cairngorms National Park, Scotland, UK, November 2013
Greylag goose (Anser anser) flying, Glenfeshie, Cairngorms National Park, Scotland, UK, April
Goldeneye (Clangula bucephala) pair on woodland loch at dawn on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
Rainbow over Scots pine (Pinus sylvestris) trees, Glen Affric, Highlands, Scotland, UK, October 2015
Patchwork of upland heather moorland and isolated Scots pine (Pinus sylvestris) trees on grouse shooting estate, Cairngorms National Park, Scotland, UK, April 2016
Osprey (Pandion haliaetus) perching on branch with fish, river and hills in background, Glenfeshie, Cairngorms National Park, Scotland, UK, August 2015
Reflections of trees in Loch Beinn a Mheadhoin, Glen Affric, Highlands, Scotland, UK, October 2015
Pine marten (Martes martes) on fallen branch in pine forest at dusk, Glenfeshie, Cairngorms National Park, Scotland, UK, April
Dense stand of Silver birch (Betula pendula), Wester Ross, Scotland, UK, February
Silver birch (Betula pendula) woodland in autumn, Craigellachie National Nature Reserve, Cairngorms National Park, Scotland, UK, October 2014
Abstract of autumnal Silver birch (Betula pendula), Cairngorms National Park, Scotland, UK, October
Scots pine (Pinus sylvestris) trees in Caledonian pine forest at sunrise, Abernethy National Nature Reserve, Cairngorms National Park, Scotland, UK, September 2014
Tangled dead boughs of veteran Scots Pine (Pinus sylvestris), Rothiemurchus Forest, Cairngorms National Park, Scotland, UK, September
View over Uath Lochans before sunrise following heavy snowfall, Cairngorms National Park, Scotland, UK, November 2015
Alt Ruadh river flowing through woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, October 2013
Sparce birch woodland in Glen Etive. Loch Etive, Highlands of Scotland, UK, January 2016
Red squirrel (Sciurus vulgaris) on branch in oak woodland with Loch Awe in background. Argyll and Bute, Scotland, UK, June
Scots Pine (Pinus sylvestris) silhouetted at dawn, Cairngorms National Park, Scotland, UK, October 2013
Willow grouse (Lagopus lagopus) Kiilopaa, Inari, Finland, January
Maple leaves backlit on lightbox, Broxwater, Cornwall, UK. November 2013
Autumnal foliage of Fireweed (Epilobium angustifolium) Kluane National Park, Yukon Territories, Canada, September
Wildebeest (Connochaetes taurinus) herd below stormy sky during migration, Masai-Mara game reserve, Kenya
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at sunrise. Kasanka National Park, Zambia
Long shadows cast by snow covered conifer trees, Kuusamo, Finland. February 2011
Sycamore tree (Acer pseudoplatanus), growing on limestone pavement, Lancashire, UK. May
Red lechwe (Kobus leche) silhouetted at sunrise, Okavango Delta, Botswana
Kokerboom or Quiver Tree (Aloe dichotoma) at night with starry sky, Kalahari, Namibia
Conifers reflected in water, Strbske Pleso, Tatra Mountains, Slovakia, June 2013
White-tailed deer (Odocoileus virginianus) in snow, Acadia National Park, Maine, USA, February
Icicles hanging from sandstone cliffs on shoreline, Apostle Islands National Lakeshore, Lake Superior, Squaw Bay, Wisconsin, February 2014
Crested Owl (Lophostrix cristata) lowland rainforests near Cristalino Jungle Lodge, Cristalino State Park, Brazil
Calcified trees at Tangled Creek in winter, Yellowstone National Park, Wyoming, USA, February 2013
Cart tracks in a flowering meadow, with Cork oaks (Quercus suber) in the background, Beja, Portugal, April
Dead trees in Deadvlei clay pan, Sossusvlei. Namib-Naukluft National Park, Namibia, September 2013
Alder (Alnus glutinosa) wood with bluebells (Hyacinthoides non-scripta) Multiple exposure. Argyll, Scotland
Herd of Impala (Aepyceros melampus) silhouetted at sunset, Ngorongoro Conservation Area, Tanzania
White-faced Capuchin (Cebus capucinus imitator) resting in palm tree. Osa PeninsulaWhite-faced Capuchin (Cebus capucinus imitator) resting in palm tree.Osa Peninsula, Costa Rica
