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Scots pine (Pinus sylvestris) silhoutte in front of Loch Maree and Slioch, Wester Ross, Scotland, UK, November 2014
Reflections in Loch Baa at dawn, Rannoch Moor, Glencoe, Lochaber, Scotland, UK, October 2014
Autumn colours and stormy clouds reflected in the calm waters of Loch Insh, Cairngorms National Park, Scotland, UK, November 2015
Zebu Cattle (Bos primigenius indicus), Los Llanos, Colombia, South America
Quaking aspen trees (Populus tremuloides) and conifers with dusting of snow, Dome Hill above Dawson City, Yukon Territories, Canada, September 2013
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
Chestnut Mustang running in snow, at ranch, Shell, Wyoming, USA. February
Great Basin Bristlecone Pine (Pinus longaeva) fallen ancient tree, White Mountains, California, USA, March
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Pine marten (Martes martes) juvenile male sitting on branch of a dead tree outside an old black woodpecker (Dryocopus martius) nest hole. Molde, Central Norway, September
Japanese fireflies (Luciola cruciata) in flight at night, endemic species, Yaku-shima UNESCO World Heritage Site, Kagoshima, Japan, May
Common juniper (Junipera communis) and Common beech (Fagus sylvatica) trees growing on pasture land grazed by livestock in summer on subalpine meadow
Underwater view of tree, with sun rays entering the water, Fibreno Lake nature reserve, Lazio, Italy
Leafless tree canopy silhouetted in winter. Gannochy, Scotland, February
Fallow deer (Dama dama) buck bellowing at dawn during the rut, Cheshire, UK, October
Monochrome image of a Hawthorn tree (Crataegus monogyna) and granite outcrop near Saddle Tor, Dartmoor National Park, Devon, England, UK, January 2011
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Rural view of rolling countryside with grazing cattle, near Frome, Somerset, UK. July 2014
American red squirrel (Tamiasciurus hudsonicus) collecting, Douglas Fir (Pseudotsuga menziesii) cones, Montana, USA
English oak tree (Quercus robur) at sunset, Monmouthshire Wales UK, March
Ash Tree (Fraxinus excelsior) growing in a field. Peak District National Park, Derbyshire, UK. August
Pampas deer (Ozotoceros bezoarticus) buck in velvet standing by flowering tree, Pantanal, Brazil
Mute Swans (Cygnus olor) foraging, southwest Finland, February
Savanna landscape wirh Grants zebras (Equus quagga boehmi) and acacia trees, Masai Mara National Reserve, Kenya, Africa
View from the top of the Ozarde waterfalls near Koyna Lake during the monsoon. Western Ghats, India, August 2010
Redpoll (Carduelis flammea) male landing on Alder (Alnus glutinosa) Surrey, England. February
Proboscis monkey (Nasalis larvatus), Sabah Malaysia, Borneo
Iberian black pig in oak woodland, Sierra de Aracena Natural Park, Huelva, Andalucia, Spain, Europe. Breed used to produce Iberico ham / Jamon Iberico
Rainforest interior views, looking up
A vine growing on Eucalyptus bark, Tompotika Peninsula, Sulawesi, Indonesia
Magnolia Rustica rubra in flower in garden, UK, May
Black bears (Ursus americanus) three cubs, one climbing, Minnesota, USA, June
Brown Bear Cub (Ursus arctos) standing behind tree. Finland, Europe, June
Aerial view of Kapok / Ceiba tree (Ceiba pentandra) in the Amazonian canopy, Yasuni National Park, Orellana, Ecuador
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
Tengmalms Owl, Boreal Owl (Aegolius funereus) looking out from its nest hole, Norway
Long eared owl (Asio otus) at night, perched on oak tree snag. Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany, August
English oak (Quercus robur) acorn. Cambridgeshire, UK. September
Fly (Mycophaga testacea) laying eggs on Porcelain fungus (Oudemansiella mucida) growing on dead Beech tree (Fagus sylvatica). Peak District National Park, Derbyshire, UK. October
Stormy light over fields near Longnor, Peak District National Park, Staffordshire, May 2014
RF - Red Squirrel (Sciurus vulgaris) adult in summer coat leaping onto fallen log. Scotland, UK. September
Robin (Erithacus rubecula) portrait on (Pinus sp), Black Isle, Scotland, UK March
Female Kestrel (Falco tinnunculus) perched in the entrance to a nest hole in an old tree, Hertfordshire, England, UK, June
American Bison (Bison bison) in habitat, Grand Teton National Park, Wyoming, USA, October
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Tropical forest, Andasibe-Mantadia National Park, Madagascar, March 2005
Detail of palm leaf, backlit in sunshine, New Caledonia
Peacock butterfly (Inachis io) feeding on Goat Willow catkins (Salix caprea), an important food source for pollinators in early spring. Peak District National Park, UK. April