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Human footprints in the sand, Sandymouth bay, Cornwall, UK
Path through Heather flowering on lowland heathland, Rockford Common, Linwood, New Forest National Park, Hampshire, England, UK, dawn, August 2011
Red fox (Vulpes vulpes) tracks in fresh snow, Jura, Switzerland
Polar bear footprint in snow {Ursus maritimus} Churchill, Canada
Polar bear (Ursus martimus) footprints in snow, Wrangel Island, Far Eastern Russia, March
Wader footprints in mudflats, Morecambe Bay, Cumbria, England, UK, February
Golden Retriever x Poodle F1b Goldendoodle puppy on paw print background
Wolf (Canis lupus) male on rocky snow covered slopes. Ulley Valley in the Himalayas, Ladakh, India
Gemsbok (Oryx gazella) aerial view of herd on sand, Namib Desert, Namibia
Peringueys adder (Bitis peringueyi) sidewinding in Namib desert, Namibia
Aerial view of cattle tracks in a salt pan, Anna Creek Station, South Australia. March, 2022
Grey wolf (Canis lupus) pack resting on a snow ledge, Hayden Valley, Yellowstone National Park, USA. January
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Trails left on sand-covered rock from Edible periwinkle (Littorina littorea) grazing on algae. Cornwall, UK. May
Firefly (Lamprohiza splendidula) light trails in woodland at night, multiple exposure. The Netherlands. June
Aerial View showing Hippopotamus trails in the swamps of the Okavango Delta, Botswana
Footprints of Arctic hare (Lepus arcticus) with Northern lights (Aurora borealis) over Senja, Norway. February
Wolverine (Gulo gulo) crosses snow field covered with animal tracks, Kamchatka, Far East Russia, April 2008
Tetrapod tracks preserved in rock at the Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Scots pine (Pinus sylvestis) cone that been partially chewed / eaten by a Red squirrel (Sciurus vulgaris) on lichen covered forest floor. Cairngorms National Park, Highlands, Scotland, UK, August
Red Squirrel (Sciurus vulgaris) foot prints in snow, Cairngorms National Park, Highlands, Scotland, UK, January
Oystercatcher (Haematopus ostralegus) footprints on sand at low tide, Norfolk, England, UK, November
Horned adder (Bitis caudalis) Swakopmund, Erongo, Namibia
Ash trees (Fraxinus excelsior) in fields next to farm track, Birchover, Peak District National Park, Derbyshire, UK, July
Sand Viper (Cerastes vipera) side winding across sand dunes, near Chingueti, Mauritania
Footprints of an Amur / Siberian tiger (Panthera tigris altaica) in sand along the edge of the Sea of Japan, Lazovskiy Zapovednik Nature Reserve, Primorsky Krai, Russian Far East. February 1992
Leatherback Turtle (Dermochelys coriacea) returning to the sea after egg laying. Cayenne, French Guiana, June
Theropod dinosaur tracks preserved in Lower Jurassic (200 million years old) rock. Near Tuba City, Arizona, USA
The road to Ojo de Perdiz, high on the altiplano, Bolivia, December 2009
Aerial view of massive arribada of Olive ridley turtle (Lepidochelys olivacea), with over 300, 000 females coming ashore to nest on 3 km of 15 km beach over period of three days and nights
Olive ridley turtle (Lepidochelys olivacea) newborn hatchling arriving at Pacific ocean during arribada, mass nesting event. Pacific coast, Oaxaca state, Mexico. November
Feeding trails from Common snails (Helix aspersa) grazing algae growing on black paint. The rows of chitinous, recurved teeth (the cuticula, part of the radula) leave a distinctive zigzag pattern
Moths (Lepidoptera) swarming with flight trails, attracted by light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands
Moths (Lepidoptera) with flight trails, attracted to light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands. June
Rear view of male Bengal tiger walking {Panthera tigris tigris} Kanha NP, India - Winner in Wildlife Photographer of Year competition
Leopard (Panthera pardus) low angle view of adult walking along track, Okavango Delta, Botswana
Common gull (Larus canus) tracks on Luskentyre Banks, South Harris Island, Outer Hebrides, Scotland, UK, June 2009
Lugworm (Anericola marina) casts on sand, Japsand, Schleswig-Holstein Wadden Sea National Park, Germany, April 2009
Mountain hare (Lepus timidus) footprints in snow, Creag Meagaidh National Nature Reserve, Scotland, UK, December 2012
European river otter (Lutra lutra) footprints crossing frozen Loch Lurgainn at dawn, with Sgorr Tuath & Sgorr Deas (Beinn an Eoin) behind, Coigach, Wester Ross, Scotland, UK, December 2010
Rock ptarmigan (Lagopus mutus) tracks in snow in winter landscape, Cairngorms NP, Highlands, Scotland, UK, February 2010
Rock ptarmigan (Lagopus mutus) tracks in snow, with low winter sun, Lochain Mountain, Cairngorms NP, Highlands, Scotland, UK, February 2010. 2020VISION Book Plate
Rock ptarmigan (Lagopus mutus) flying from boulder field in winter plumage, Cairngorms NP, Highlands, Scotland, UK, February
Peringueys adder / Sidewinding adder (Bitis peringueyi), sidewinding, Namib Desert, Namibia, May
A typical European Grey wolf (Canis lupus) track in the snow, Val Borbera, Liguria, Italy
Wood pigeon (Columba palumbus) footprints in snow, Norfolk, England, March
