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Santa Cruz Galapagos tortoise (Chelonoidis nigra porteri) feeding on grass, Santa Cruz Highlands, Galapagos
Floreana giant tortoise hybrid descendant (Chelonoidis elephantopus), Fausto Llerena Giant Trotoise Breeding Centre, Galapagos National park Directorate, Puerto Ayora, Santa Cruz Island, Galapagos
Alcedo giant tortoises (Chelonoidis vandenburghi) group resting in water to keep cool, Alcedo Volcano, Isabela Island, Galapagos
Western Santa Cruz giant tortoise (Chelonoidis porteri) portrait, Highlands, Santa Cruz Island, Galapagos
Volcan Alcedo giant tortoises (Chelonoidis nigra vandenburghi) among steaming fumaroles on western caldera rim, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Wolf giant tortoise (Chelonoidis becki) group drinking from small puddles formed by fine drizzle. Wolf Volcano, Isabela Island, Galapagos
Wolf giant tortoise (Chelonoidis becki) portrait. Wolf Volcano, Isabela Island, Galapagos
Wolf giant tortoise (Chelonoidis becki) in habitat. Hybrids of mixed parentage with different shell shapes are scattered far and wide on the rugged west slope of Wolf Volcano
White-cheeked pintails (Anas bahamensis) with Western Santa Cruz tortoise (Chelonoidis porteri) at pond, Santa Cruz, Galapagos
Galapagos dove (Zenadia galapagensis) on head of Galapagos giant tortoise, (Chelonoidis darwini) Santiago Highlands, Galapagos
Espanola saddelback tortoise (Chelonoidis hoodensis) pair mating, Espanola Island, Galapagos
Leopard / mountain tortoise (Stigmochelys pardalis) with elephant bull (Loxodonta africana) behind in water, Addo elephant national park, Eastern Cape, South Africa, September
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Aldabra Giant Tortoises (Aldabrachelys gigantea) resting in a pool to keep cool, Grand Terre, Natural World Heritage Site, Aldabra
Floreana giant tortoise hybrid descendants (Chelonoidis elephantopus) feeding on flower. These are descendants of the extinct Floreana giant tortoise, but are hybridised with other species
Hermanns / Dalmatian tortoise (Testudo hermanni hercegovinensis) near Svitava Lake, Hutovo Blato Nature Park, Bosnia and Herzegovina, May 2009
Yellow footed tortoise (Chelonoidis denticulata) portrait, Yasuni National Park, Orellana, Ecuador. Vulnerable species
Galapagos giant tortoise (Chelonoidis nigra) portrait, Santa Cruz Island, Galapagos Islands, Ecuador. Endangered
Red-footed tortoise (Chelonoidis carbonaria) portrait, Pantanal wetlands, Mato Grosso do Sul, Brazil
Santa Cruz giant-tortoise (Chelonoidis porteri) covered in mud, Santa Cruz Island, Galapagos National Park, Galapagos Isalnds. Critically endangered
Group of Santa Cruz giant-tortoises (Chelonoidis porteri) wallowing in mud, Santa Cruz Island, Galapagos National Park, Galapagos Islands. Critically endangered
Fernandina giant-tortoise (Chelonoidis phantasticus) head portrait, Fernandina Island, Galapagos National Park, Galapagos Islands. Critically endangered
Detail of hind toes of Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Darwin's Small ground finch (Geospiza fuliginosa) cleaning Alcedo giant tortoise (Chelonoidis vandenburghi) of ticks, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Old male Espanola giant Galapagos tortoise (Chelonoidis hoodensis) stretching its neck in clearing, Las Tunas, Espanola Island, Galapagos Islands, Ecuador
Eastern Hermann's tortoise (Testudo hermanni boettgeri) hatching from egg, mouth open. Captive, occurs in South East Europe
Sierra Negra giant tortoise (Chelonoidis guntheri) walking, Isabela Island, Galapagos, South America
Pair of Leopard tortoises (Stigmochelys pardalis) mating, Karoo, South Africa
Alcedo giant tortoise (Chelonoidis vandenburghi), Alcedo Volcano, Isabela Island. This is where the largest extant population exists
Espanola giant tortoise (Chelonoidis hoodensis), in grasses, Santa Fe Island, Galapagos
Alcedo giant tortoises (Chelonoidis vandenburghi) and volcanic landscape with fumeroles, Alcedo Volcano, Isabela Island, Galapagos
Epanola saddelback tortoise (Chelonoidis hoodensis), with Galapagos mockingbird (Mimus parvulus) perched on carapace, Espnola Island, Galapagos
Alcedo Galapagos tortoise (Chelonoidis nigra vandenburghi) group wallowing in mud, Alcedo Volcan, Isabela Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi), group bathing. Alcedo Volcano, Isabela Island, Galapagos. 2017
Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp)Wolf Volcano giant tortoise (Chelonoidis nigra becki) portrait with ground finch (Geospiza sp. ) sitting on foot, Isabela Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in pool, Alcedo Volcano, Isabela Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) silhouetted at sunrise, Alcedo Volcano, Isabela Island, Galapagos
Alcedo giant tortoise (Chelonoidis vandenburghi) group resting in water at night, Alcedo Volcano, Isabela Island, Galapagos
Hermanns tortoise (Testudo hermanni) Shebeniku-Jabllanica National Park, Albania, June 2009
Hermanns tortoise (Testudo hermanni) in a meadow, Patras area, The Peloponnese, Greece, May 2009
Close-up of Hermanns tortoise (Testudo hermanni) head, Djerdap National Park, Serbia, June 2009
Hermanns tortoise (Testudo hermanni) portrait, Djerdap National Park, Serbia, June 2009
Greek land tortoise (Testudo graeca ibera) head portrait, Lake Kerkini, Macedonia, Greece, May 2009
Greek tortoise (Testudo graeca ibera) portrait, Lake Kerkini, Macedonia, Greece, May 2009
Greek land / Spur thighed tortoise (Testudo graeca) head portrait, Lake Kerkini, Macedonia, Greece, May 2009 WWE BOOK
