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Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
African Wild Dog (Lycaon pictus) pack passing infront of large African elephant (Loxodonta africana) Mana Pools National Park, Zimbabwe October 2012
Ogre faced / Net-casting spider {Deinopis sp} with web held between legs that it will stretch over prey that walks below it. Masoala Peninsula National Park, north east Madagascar
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Silhouette of Barn Swallow (Hirundo rustica) flying over water, hawking for insects, Berwickshire, Scotland, UK, July
Portrait of a Common Wasp (Vespula vulgaris). UK. Focus stacked image (dead specimen)
Boomslang (Dispholidus typus) Immature male hunting in thorn bush above deHoop Vlei, Western Cape, South Africa, December
Mozambique Spitting Cobra (Naja mossambica) ejecting venom Kruger, South Africa, Controlled conditions
Male Adder (Vipera berus) tasting the air. Cannock Chase, Staffordshire, UK, October
Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean
Dalmatian pelican (Pelecanus crispus) low angle perspective of open bill, Lake Kerkini, Greece, February. Winner, Eric Hosking award, 2011 Wildlife Photographer of the Year competition
Lemon shark (Negaprion brevirostris) under sun rays. Little Bahama Bank, Bahamas. West Atlantic Ocean
A large female Tiger Shark (Galeocerdo cuvier) swimming in open water. Grand Bahama, Bahamas. Tropical West Atlantic Ocean. March
Sea holly (Eryngium maritimum) growing in sand dune, Menorca, Balearic Islands, Spain, Europe
Big close up of the eyes of a European Brown bear (Ursus arctos) captive
Angler fish (Melanocetus murrayi) Mid-Atlantic Ridge, North Atlantic Ocean
Mandrill (Mandrillus sphinx) male showing aggression, captive
Brown Hare (Lepus europaeus) close-up of fur and eye. Derbyshire, UK, March
Indian moon / Indian luna Moth (Actias selene) close up of defensive spines on caterpillar. Captive
Alpine ibex (Capra ibex) adult male chasing two young males away, Gran Paradiso National Park, Italy, July
Ivory Ornamental Tarantulas (Poecilotheria subfusca) from Sri Lanka
Costa Rican Redleg Tarantula (Megaphobema mesomelas), captive from South America
Portrait of Waglers / Temple Pitviper (Tropidolaemus wagleri) showing the thermo-receptive pit organ between the eye and mouth. Controlled conditions. Mindanao, Philippines, February
Large adult male Leopard (Panthera pardus) walking through the bush at night, Sabi Sand Private Game Reserve, South Africa, June
Great white shark (Carcharodon carcharias) portrait, Guadalupe Island, Mexico, Pacific Ocean
Head portrait of Sumatran tiger (Panthera tigris sumatrae) with face half cast in shadow, with aggressive expression, captive
Maine Coon female cat, Serafin, in fierce defensive posture
Smoke from bush fire in Pilbara region, Western Australia. August 2009
Great skua (Stercocarius skua) in flight over Gannet (Morus bassanus) colony, looking for small chicks, whilst Gannets call defensively, Shetland Isles, July
Large / Pot Bellied Seahorse (Hippocampus abdominalis) face portrait. Blairgowrie Marina, Mornington Peninsular, Victoria, Australia, March
Two Red Foxes (Vulpes vulpes) fighting. The Netherlands. August
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Goldfinches (Carduelis carduelis) squabbling near seed feeder, Berwickshire, Scotland, April
Ruffs displaying and fighting (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Arctic tern (Sterna paradisaea) adult hovering directly in front of the sun, defending nest from perceived threat, Farne Islands, UK. June Non-ex
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
Alpine Chough / Yellow-billed Chough (Pyrrhocorax graculus) flock in flight against mountains, Dolomites, Italy. Digital composite
Western Diamondback Rattlesnake (Crotalus atrox), adult in striking pose in wildflower field. Laredo, Webb County, South Texas, USA, April
Close-up portrait of a Red Fox (Vulpes vulpes). Vosges, France, June
Narwhal (Monodon monoceros) showing tusk above water surface. Baffin Island, Nunavut, Canada, April
White-lipped / Green tree viper (Cryptelytrops albolabris) captive, from SE Asia
Dramatic seas crashing onto a rocky shore. The Broken Group Islands from The Wild Pacific Trail, Ucluelet, West coast of Vancouver Island, Canada, September 2010
Caribbean reef shark (Carcharhinus perezi) over sand ripples, Walkers Cay, Bahamas. Tropical West Atlantic Ocean. Taken in natural light
Catesbys snail-eating snake (Dipsas catesbyi) on Heliconia plant, Pacaya Samiria National Reserve, Amazon Basin, Loreto, Peru
Lemon shark (Negaprion brevirostris) in shallow water with reflection at the surface. Little Bahama Bank, Bahamas
A lone Caribbean reef shark (Carcharhinus perezi) cruises over sand ripples. Walkers Cay, Northern Bahama Islands, Republic of Bahamas
Lighthouse on Snafellsness Peninsula, Iceland, July 2008