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Young urban Red fox (Vulpes vulpes). Bristol, UK. August
Dalmatian Pelican (Pelecanus crispus) portrait, close-up of eyes, Lake Kerkini, Greece. February
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
RF- Banded demoiselle (Calopteryx splendens), resting on dew covered reed, Lower Tamar Lakes, Cornwall/Devon border, UK. May
Golden Eagle (Aquila chrysaetos) in flight rear view, Kuusamo, Finland, February
Great white shark (Carcharodon carcharias) from above. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
RF - Eyelash Pit Viper (Bothriechis schlegelii) Costa Rica
Ribbon sawtail fish (Idiacanthus fasciola) from Atlantic Ocean, at a depth of 800-1000m
Jackdaw (Corvus monedula) pair chasing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel and its mate
Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Fairy Pools and Black Cuillins, Glen Brittle, Isle of Skye, Inner Hebrides, Scotland, UK. February 2017
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
RF- Black rhinoceros (Diceros bicornis) looking threatening, Etosha National Park, Namibia, June. Endangered species
Great white shark (Carcharodon carcharias) Guadalupe Island or Isla Guadalupe, Pacific Ocean, Mexico, September. Vulnerable species
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Puff Adder (Bitis arietans) captive occurs in Tanzania
African horned viper (Cerastes cerastes), captive
Yacare Caiman (Caiman yacare) with mouth open to keep cool, Pantanal, Brazil
Marsh harrier (Circus aeruginosus) adult female circling over Oilseed rape crop, Norfolk, England, May. Digital composite
White tailed eagle (Haliaeetus albicilla) in flight with snowy background, Rausu, Japan, February
Yellow lipped sea snake (Laticauda colubrina) dives back down to the reef after surfacing for breath of air. Apo Island, Dumaguete, Negros, Philippines. Bohol Sea, Tropical West Pacific Ocean
Stag beetle (Lucanus cervus) silhouetted against oak tree leaf. Elbe, Germany, June
Cup fungus (Cookeina sp) Danum Valley, Sabah, Borneo
Great white shark (Carcharodon carcharias) swimming close to the surface, Guadalupe Island, Baja California, Mexico, Pacific Ocean
Great white shark (Carcharodon carcharias) swimming, Guadalupe Island, Mexico, Pacific Ocean
Jackdaw (Corvus monedula) pair dive bombing and harassing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel
Jackdaw (Corvus monedula) swooping down calling with claws raised to threaten a Grey squirrel (Sciurus carolinensis) in the entrance to a nest box the bird wants to nest in which is already occupied
Jackdaw (Corvus monedula) swooping in very close to threaten a Grey squirrel (Sciurus carolinensis) as it stands in the entrance to a nest box the bird wants to nest in which is already occupied by
Texas brown tarantula (Aphonopelma hentzi) in defensive posture, Comanche National Grasslands, Colorado, USA
Striped Slender Robberfly (Leptogaster cylindrica) Ledston, Yorkshire, July. Focus stacked image
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
RF - Herring gull (Larus argentatus) with blue sky, England, UK, Digital composite
Polar bear (Ursus maritimus) young bear in freezing water during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska, Beaufort Sea
Lioness (Panthera leo) and Thomsons gazelles (Eudorcas thomsonii) silhouetted at sunset, Masai-Mara game reserve, Kenya
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
Komodo Dragon (Varanus komodoensis) close up, Komodo National Park, Indonesia
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America
Leopard (Panthera pardus) portrait, Samburu National Reserve, Kenya
Black rhino (Diceros bicornis) looking threatening, Nakuru National Park, Kenya
White rhino (Ceratotherium simum) in the rain, Nakuru National Park, Kenya
Large female Southern stingray (Hypanus americanus) in shallow water, under storm clouds, Stingray City, Grand Cayman. Stingray City is feeding site for stingrays and bait was used on this shoot
Orcas (Orcinus orca) pod feeding on herring, wide shot showing surrounding landscape, Iceland, January 2013