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Kakapo (Strigops habroptilus) close up showing sensory facial feathers, Codfish Island / Whenua Hou, Southland, New Zealand, February. Critically endangered
RF - Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Silhouetted Dartmoor pony (Equus caballus) at sunrise, Combestone Tor, Dartmoor National Park, Devon, England, UK, January
European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June
Close up face portrait of male Orang Utan (Pongo pygmaeus) Captive, Netherlands
Female chimpanzee (Pan troglodytes troglodytes) portrait. Yoko, age 35 years, is the dominant female of the group, Conkouati-Douli National Park, Republic of Congo, Africa
RF- Head portrait of Bulldog puppy with chin on paws
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
African lion (Panthera leo), resting on a rock, surrounded by flies, Masai Mara Game Reserve, Kenya. November
RF- African Lion (Panthera leo) young male at sunrise, Etosha National Park, Namibia
Jaguar (Panthera onca) portrait, lying down, captive
Whooper Swan (Cygnus cygnus) standing in snowfall, Hokkaido, Japan, February
European hornet (Vespa crabro) worker in flight, Surrey, England, August
RF- Pair of Red Sea anemonefish (Amphiprion bicinctus) in Magnificent sea anemone (Heteractis magnifica), which has closed up in the late afternoon revealing its red skirt. St Johns Reef. Egypt
Kelp gull (Larus dominicanus) on rocky shore with crashing waves at sunset, Bird Island, Lamberts Bay, Western Cape province, South Africa, September 2012
Indochinese tiger (Panthera tigris corbetti) looking up, behind screen of leaves, captive, occurs in South East Asia
RF- Small jellyfish (Pelagia noctiluca) swimming beneath surface. Marine Protected Area of Portofino (Area Marina Protetta, Portofino), Liguria, Italy. Mediterranean Sea
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
RF- Siberian tiger (Panthera tigris altaica) young cub, lying down, looking away, captive
Mountain Gorilla (Gorilla gorilla beringei) mother holding baby twins age five months, Parc National des Volcans, Rwanda. Endangered species
Harvest mice (Micromys minutus), captive, UK, June
Mountain Hare (Lepus timidus) side view portrait, Cairngorms, Scotland, February
Snow Monkey (Macaca fuscata) with snow covered paw in front of mouth, Nagano, Japan, February
Barred owl (Strix varia) stretching wings, Corkscrew Swamp Audubon Sanctuary, Florida, USA, March
RF - Bluebell (Hyacinthoides non-scripta) close-up. Lanhydrock woodland, Cornwall, UK. May
Dalmatian Pelicans (Pelecanus crispus) portrait. Lake Kerkini, Greece, March 2012
Red Fox (Vulpes vulpes) in wide landscape with Kronotsky Volcano on the horizon. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska