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Territories Prints
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Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek
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Two Black woodpeckers (Dryocopus martius) males on tree trunk in territorial dispute
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RF - Frill-necked dragon (Chlamydosaurus kingii)
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Northern gannet (Morus bassanus)
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Northern gannet (Morus bassanus)
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Northern gannet (Morus bassanus)
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Northern gannet (Morus bassanus)
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Two Black-necked stork (Ephippiorhynchus asiaticus) juveniles
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Aldabra rails (Dryolimnas cuvieri aldabranus) pair calling
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European otter (Lutra lutra) urinating on a rock
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Red deer (Cervus elaphus) stag rubbing its antlers against a tree trunk
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Red deer (Cervus elaphus) stag rubbing its antlers against a tree trunk
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Fulmar (Fulmarus glacialis) perched on cliff top calling
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RF - Fulmar (Fulmarus glacialis) perched on cliff top calling
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Great skua (Stercorarius skua)
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Two Northern gannets (Morus bassanus) in flight entwined in mortal combat over the nesting colony
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Two Northern gannets (Morus bassanus) in flight entwined in mortal combat over the nesting colony
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Two Northern gannet (Morus bassanus) pairs at nesting colony in territorial dispute
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Two Puffins (Fratercula arctica) male
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RF - Mute swans (Cygnus olor) pair approaching two other swans in foreground in a territorial dispute
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Coot (Fulica atra) displaying aggression in a territorial fight
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Fallow deer (Dama dama) buck, scent marking among ferns
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Grey wolf (Canis lupus) alpha-male, scent marking it's territory, Finland. August
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Sambar (Rusa unicolor) male, rubbing its antlers against a tree
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Red fox (Vulpes vulpes) attacking a Badger (Meles meles) over a dispute linked to territory and food reserves
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Red fox (Vulpes vulpes) attacking a Badger (Meles meles) over a dispute linked to territory and food reserves
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Apennine wolf (Canis lupus italicus) patrolling its territory near an alpine skyline
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Two Giant cuttlefish (Sepia apama) males
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Freshly laid eggs of the Giant Cuttlefish (Sepia apama) that have been laid in a recess under a rocky ledge
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Impala (Aepyceros melampus) male
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Superb lyrebird (Menura novaehollandiae) male
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Lion (Panthera leo) dominant male
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Lion (Panthera leo) dominant male
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Australasian crested grebe (Podiceps cristatus australis)
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Two Speckled wood butterfly (Pararge aegeria) males chasing each other to defend territory in woodland
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Two Pied herons (Egretta picata) in breeding plumage
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Two Pied herons (Egretta picata) in breeding plumage
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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Bengal tiger (Panthera tigris tigris) standing on hind legs scratching tree trunk
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A sponge and coral reef scene with two Caesar grunts (Haemulon carbonarium) battling over territory
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Wool carder bee (Anthidium manicatum) male
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Two White-belly damselfish (Amblyglyphidodon leucogaster) male
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Northern black korhaan (Afrotis afraoides) territorial calling
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Southern red bishop (Euplectes orix) male guarding territory
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Two Striped surgeonfish (Acanthurus lineatus) jousting along a territorial boundary in the shallow water
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RF - Group of Red lionfish (Pterois volitans) engaging in cooperative hunting over coral reef
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Bengal tiger (Panthera tigris tigris) female
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Bengal tiger (Panthera tigris tigris) male
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Bengal tiger (Panthera tigris tigris) spraying urine on tree trunk to mark territory or communicate with other tigers
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Bengal tiger (Panthera tigris tigris) spraying urine on tree trunk to mark territory or communicate with other tigers
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Bengal tiger (Panthera tigris tigris) female with three cubs
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Ethiopian wolves (Canis simensis) chasing an intruder out of their territory
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Two Ethiopian wolves (Canis simensis) patrolling their territory boundary
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Ethiopian wolf (Canis simensis) pack members patrolling their territory borders
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Ethiopian wolf (Canis simensis) pack members patrolling their territory borders
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Ethiopian wolves (Canis simensis) patrolling their territory boundary
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Two Ethiopian wolves (Canis simensis) patrolling their territory boundary
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Ethiopian wolf (Canis simensis) patrolling the pack's territory
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RF - Frill-necked dragon (Chlamydosaurus kingii) male
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Two Shelduck (Tadorna tadorna) drakes chasing each other on water during territorial dispute
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Two Masai ostriches (Struthio camelus massaicus) mayor female and satellite female
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Two Masai ostriches (Struthio camelus massaicus) mayor female and satellite female
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Three Masai ostriches (Struthio camelus massaicus) mayor female and satellite females
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Three Masai ostriches (Struthio camelus massaicus) mayor female and satellite females
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Three Masai ostriches (Struthio camelus massaicus) mayor female and satellite females
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Three Masai ostriches (Struthio camelus massaicus) mayor female and satellite females
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Two Masai ostriches (Struthio camelus massaicus) mayor female and satellite female
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Two Musk ox (Ovibos moschatus) bulls displaying aggression
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Arctic skua (Stercorarius parasiticus) female
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Two Musk ox (Ovibos moschatus) bulls sparring
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Two Musk ox (Ovibos moschatus) bulls sparring
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Frill-necked dragon (Chlamydosaurus kingii) male
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Dunnock (Prunella modularis) perched on branch performing wing flapping display
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Dunnock (Prunella modularis) perched on branch performing wing flapping display
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Dunnock (Prunella modularis) perched on branch performing wing flapping display
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Dunnock (Prunella modularis) perched on branch performing wing flapping display
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Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it
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Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it
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Lion (Panthera leo) male, scent marking its territory at dawn
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