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European river otter (Lutra lutra) swimming over kelp, Shetland, Scotland, UK, April
Killer whale (Orcinus orca) tail slapping at sunset, transient race, Vancouver Island, British Columbia, Canada, July
Pool Frog (Pelophylax lessonae) split level view, near Crisan village, Danube Delta, Romania, June
Bottlenose Dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal
Candy stripe shrimp (Lebbeus grandimanus) on a yellow sponge. Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
A portrait of a large male Dusky grouper (Epinephelus marginatus). Lavezzi Islands, between Sardinia and Corsica in the Mediterranean Sea
Tail of Southern right whale (Eubalaena australis) Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean, October
Barrel Jellyfish (Rhizostoma pulmo) against crepuscular light rays. Sark, British Channel Islands, July
Sequence of a breaching Southern right whale (Eubalaena australis) Golfo Nuevo, Peninsula Valdes, UNESCO Natural World Heritage Site, Chubut, Patagonia, Argentina, Atlantic Ocean
Amphipod (Iphimedia obesa) on top of a colony of Deadmans fingers (Alcyonium digitatum) coral, Loch Carron, Ross and Cromarty, Scotland, UK, April
A portrait of a Mediterranean anthias (Anthias anthias) against open water. Secca Del Papa, Tavolara Marine Protected Area, Sardinia, Italy. Mediterranean Sea
European river otter (Lutra lutra) swimming in the sea, Isle of Mull, Inner Hebrides, Scotland, UK, December
Blue shark (Prionace glauca) female off the coast of Portugal. August
Close up detail photo of a Spotted moray (Gymnothorax isingteena), Baa Atoll, Maldives. Indian Ocean
A pair of Purple sea stars (Pisaster ochraceus) climb on anemones in shallow water, beneath trees in Browning Pass, Port Hardy, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Aggregation of Spider crabs (Maja squinado) in shallow water off Burton Bradstock, Dorset, UK, August. 2020VISION Book Plate
Eye detail of Topknot (Zeugopterus punctatus) flatfish, Loch Fyne, Argyll, Scotland, UK, July
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
Group of Jewel anemones (Corynactis viridis) photographed under blue light to show their fluorescence, Plymouth Sound, Devon, UK, July
Portrait of a Grunt sculpin (Rhamphocottus richardsonii) yawns as it moves through soft corals and sponges. This unusual looking fish is evolved to resemble the giant acorn barnacle
Eye detail of Red Irish lord (Hemilepidotus hemilepidotus). Browning Pass, Port Hardy, Vancouver Island, Canada
Pacific giant octopus (Enteroctopus dofleini) hunts for food on a colourful reef in Browning Pass, Vancouver Island, British Columbia, Canada
Grey seal (Halichoerus grypus) surfacing beneath cliffs, Lundy Island, Devon, UK, June
A pair of captive Common / Short snouted seahorses (Hippocampus hippocampus) in aquarium, Weymouth Sealife Centre, Dorset, England, UK
Red gorgonians (Lophogorgia chilensis) growing beneath a Giant kelp (Macrocystis pyrifera) forest, with female California sheephead (Semicossyphus pulcher). Santa Barbara Island, Channel Islands
Amphipod travelling on sea gooseberry (Ctenophora) Browning Pass, Vancouver Island, British Columbia, Canada. North East Pacific Ocean
Portrait of a male River blenny (Salaria fluviatilis) River Flumendosa, Gennargentu National Park, Sardinia, Italy
Grey seal (Halichoerus grypus) young seal silhouetted against the sun as it dives after breathing, Lundy Island, Devon, UK, Bristol Channel, August
Grey seal (Halichoerus grypus) split level view of young investigating camera lens, Lundy Island, Devon, UK, Bristol Channel, August
Harbour seal (Phoca vitulina) in the kelp forest (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean
California sea lion (Zalophus californianus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath Eureka oil rig, Los Angeles, California, USA, Pacific Ocean
Florida manatee (Trichechus manatus latirostrus) being cleaned by blue gill sunfish (Lepomis macrochirus). Three Sisters Spring, Crystal River, Florida. USA. February 2010
Grey seal (Halichoerus grypus) portrait underwater, Farne Islands, Northumberland, England, UK, August
Underwater landscape in peat pond, Fribourg, Switzerland, August
Jewel Anemones (Corynactis haddoni) Poor Knights Islands, New Zealand, February
Yellow zoanthids (Parazoanthus sp. ) Poor Knights Islands, New Zealand, JanuaryYellow zoanthids (Parazoanthus sp.) Poor Knights Islands, New Zealand, January
Yellow cave coral {Leptosammia pruvoti} off Sark, Channel Isles, UK
A blue light fluorescence photo of Jewel anemones (Corynactis viridis). The blue light source excites the fluorescent pigments in the anemones, which emit light of other colours
Light-bulb sea squirts (Clavelina lepadiformis), a colonial filter feeding invertebrate, fluoresce green when photographed under deep blue light. Shetland Islands, Scotland, UK, June
Waikoropupu Springs, Golden Bay, South Island, New Zealand
Three hairy sea spiders (Tanystylum grossifemorum) feeding on red soft coral (Gersemia rubiformis). Browning Pass, Port Hardy, Vancouver Island. British Columbia. Canada. North East Pacific Ocean
Young grey seal (Halichoerus grypus) lying on sand, Helgoland, Germany
Aggregation of Spider crabs (Maja squinado) in shallow water off Burton Bradstock, Dorset, UK, August
Whale sharks (Rhincodon typus) bottle feeding, by floating stationary, upright in the water and gulping in food, in the plankton rich, green waters of La Paz bay. La Paz, Baja California Sur, Mexico
Bottlenose dolphin (Tursiops truncatus) playing in the waves of an oil tanker, Port Aransas, Texas, USA
Split image of Blue shark (Prionace glauca) close to the surface, Pico Island, Azores, Portugal, Atlantic Ocean, July
Portrait of a Yellow seahorse (Hippocampus guttulatus), Capo Galera, Alghero, Sardinia, Italy, Mediterranean Sea
Brown trout (Salmo trutta) resting on river bed near the Aigas Field Studies Centre, Inverness-shire, Scotland, July