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RF - Dapple grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
Orangutan baby (Pongo pygmaeus) head portrait of baby, Semengoh Nature reserve, Sarawak, Borneo, Malaysia. Endangered
Rose-ringed / ring-necked parakeets (Psittacula krameri) in flight on way to roost in an urban cemetery, London, UK. January
Portrait of a young Little Owl (Athene noctua)
Rough coated Jack Russell Terrier puppy, black and tan, portrait
Little owl (Athene noctua) Sado Estuary, Portugal. June
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Yellow mongoose (Cynictis penicillata) standing on hind legs with young, Kgalagadi Transfrontier Park, South Africa, February
Yellow Guinea pig standing up and squeaking, against white background
Indri (Indri indri) calling, Andasibe-Mantadia NP, Madagascar
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Startled Eurasian lynx (Lynx lynx) kitten, aged eight weeks, breeding and reintroduction program, Germany. Captive. August
Great bittern (Botaurus stellaris) head and neck, lookin up through reeds. Suffolk, UK. November
Smooth trunkfish (Lactophrys triqueter), portrait. Bonaire, Dutch Caribbean
Lion (Panthera leo) male with scars photographed with side-lit spot light at night. Greater Kruger National Park, South Africa, April
Portrait of rabbit in leaf litter, Okunoshima Rabbit Island, Takehara, Hiroshima, Japan
Peringueys / Sidewinding adder (Bitis peringueyi) hiding in shallow sand, Namib Desert, Namibia. This little endemic snake is one of the smallest adders in the world
Bighorn Sheep (Ovis canadensis) Yellowstone National Park, Wyoming, USA. January
Snowy egret (Egretta thula) with plumes erect, St. Augustine, Florida, USA, April
Horned puffin (Fratercula corniculata) close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July
Spotted hyena (Crocuta crocuta) hunting Lesser flamingoes (Phoenicopterus minor) Lake Nakuru, Kenya
Capercaillie (Tetrao urogallus) male in flight, Jalasjarvi, Finland, April
Male Yellow boxfish (Ostracion cubicus) displaying, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Adult Western / Horsfields tarsier (Tarsius bancanus) in forest understorey at night. Lowland dipterocarp rainforest, Danum Valley, Sabah, Borneo
Black Lemur (Eulemur macaco) male, Nosy Komba, Madagascar
Wild Cat (Felis silvestris) in mist. Vosges, France, August
Long-eared owl (Asio otus) perched in tree amongst autumn leaves, Czech Republic, October. Captive
RF- Eurasian Eagle owl (Bubo bubo) adult perched on rocky outcrop with the Super Full Moon, September 28th 2015, Southern Norway. August. Multiple exposure
RF- Wide angle portrait of Hairy frogfish (Antennarius striatus) waiting to ambush prey on black sand slope. Lembeh Strait, North Sulawesi, Indonesia. Molucca Sea
Asiatic lion cub (Panthera leo persica) looking up into tree, possibly at a bird, Gir Forest NP, Gujarat, India
Galapagos flycatcher (Myiarchus magnirostris) in flight, Santiago Island, Galapagos National Park, Galapagos Islands
Black Welsh mountain sheep lamb calling. Wales UK, March
White-eyed moray (Muraena thyrsoides) close up head portrait, Bali, Indonesia, Indo-Pacific
Common pheasant (Phasianus colchicus) male flapping wings and calling, Pusztaszer reserve, Hungary. May
Gannet (Morus bassanus) chick with fluffy with mouth open. Shetland Islands, Scotland, UK, August
Demoiselle cranes (Anthropoides virgo) flock, India
RF - Great horned owl (Bubo virginianus) in woodland, captive; Alaska, USA
RF- Tengmalms owl (Aegolius funereus) perched in tree, Bergslagen, Sweden, June 2009
Tabby kitten, Picasso, age 10 weeks, leaping out of wicker basket with the word Tadaa! added digitally underneath
Close up of tiny (10mm) Pygmy seahorse (Hippocampus bargibanti) living disguised in Muricella sp. sea fan. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Infant Golden-crowned Sifaka (Propithecus tattersalli) on its mothers back. Forests adjacent to the village of Andranotsimaty, near Daraina, northern Madagascar. Critically endangered species
Lurcher dog, Kite, lying with head up, against white background
Black-and-white tuxedo male kitten, Tuxie, 8 weeks, with black-and-white Dutch rabbit, against white background
Hare, (Lepus europaeus) jumping in snowy field, Vosges, France, February
Male Little bustard (Tetrax tetrax) displaying, Catalonia, Spain, May
Andean Gull (Chroicocephalus serranus) calling in flight, Cotopaxi, Ecuador