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Ray of light shining through thunder clouds over Lake Ohrid, Galicica National Park, Macedonia, June 2009
Underwater view of tree, with sun rays entering the water, Fibreno Lake nature reserve, Lazio, Italy
King penguin (Aptenodytes patagonicus) chicks in mass breeding colony, St Andrews Bay, South Georgia. Taken on location for BBC Frozen Planet series, 2008
Sun reflecting on sea surface with rocks on beach, Dail Beag Beach, Lewis, Outer Hebrides, Scotland, UK, June 2009
Sun shining through clouds over Lake Skadar near Murici, Lake Skadar National Park, Montenegro, May 2008
Sun shining through clouds, Lake Skadar, Lake Skadar National Park, Montenegro, May 2008
Ferns in a valley, Pollino National Park, Basilicata, Italy, May 2009
Sun shining through trees in Common alder (Alnus glutinosa) forest, Klampenborg Dyrehaven, Denmark, September 2008
European oak (Quercus robur) with rays of sunlight, Klampenborg Dyrehaven, Denmark, September 2008
Rays of light shining through mist surrounding Black pines (Pinus nigra) Crna Poda Natural Reserve, Tara Canyon, Durmitor NP, Montenegro, October 2008
Brown Bear (Ursus arctos) silhouetted at dawn, Suomussalmi, Kainuu region, Finland, Europe, June 2008, Exclusive Japanese calendar rights for 2014