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Kiang (Equus kiang) herd grazing on steppe in evening light, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Horses grazing in steppe grassland, Altanbulag, Mongolia
Clouds over steppe grassland, Altanbulag, Mongolia
Juvenile Pallas cat (Otocolobus manul) outside den, Mongolia, June
View of a double rainbow over steppe grassland, Mongolia, 2018
Close up of a juvenile Pallas cat (Otocolobus manul) resting in its den, Mongolia, June
Little owl (Athene noctua) taking off, Bagerova Steppe, Kerch Peninsula, Crimea, Ukraine, July 2009
Chilean flamingos (Phoenicopterus chilensis) group of three in flight, Lago Amarga, Torres del Paine National Park, Patagonia, Chile
Saiga antelope (Saiga tatarica) males drinking Astrakhan Steppe, Southern Russia
Grey wolf (Canis lupus) with Saiga antelope (Saiga tatarica) in the background, Astrakhan Steppe, Southern Russia
Male Saiga antelope (Saiga tatarica) running, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009. WWE OUTDOOR EXHIBITION. WWE BOOK
Stone curlew (Burhinus oedicnemus) Bagerova Steppe, Kerch Peninsula, Crimea, Ukraine, July 2009
Stone curlew (Burhinus oedicnemus) by nest, Bagerova Steppe, Kerch Peninsula, Crimea, Ukraine, July 2009
Little owl (Athene noctua) perched on a fence post, stretching its wings, Castro Verde, Alentejo, Portugal, April
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
European rabbit (Oryctolagus cuniculus) in steppe habitat. Alfaro, La Rioja, Spain. Endangered species
Altai marmot (Marmota baibacina) in steppe landscape, Ukok plateau, Altai Republic, Russia
Yak (Bos grunniens) herd on plain beside river, mountains in background. Near Litang, Kham, Garze Tibetan Autonomous Prefecture, Sichuan, China. October 2016
Wild yak (Bos mutus) in snow covered steppe, mountains in background. Hoh Xil Nature Reserve, Tibetan plateau, Qinghai, China. October 2019
Grey wolf (Canis lupus) Astrakhan Steppe, Southern Russia
Wild tulip (Tulipa schrenkii) flower in frost, Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Frost covered Wild tulip (Tulipa schrenkii) Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Demoiselle crane (Anthropoides virgo) tending two eggs in its nest, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Female Saiga antelope (Saiga tatarica) feeding, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Saiga antelope (Saiga tatarica) with two calves suckling near Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Frost covered Wild tulips (Tulipa schrenkii) in flower, Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Two male Saiga antelopes (Saiga tatarica) Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Three female Saiga antelopes (Saiga tatarica) Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Saiga antelope (Saiga tatarica) herd at salt lick, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Female Saiga antelope (Saiga tatarica) grazing, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009. WWE INDOOR EXHIBITION
Saiga antelope (Saiga tatarica) herd silhouetted, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Male Saiga antelope (Saiga tatarica) silhouetted, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Newborn Saiga antelope (Saiga tatarica) lying in grass, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Steppe eagle (Aquila nipalensis) on nest with chicks, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Two Steppe eagle (Aquila nipalensis) chicks, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Demoiselle crane (Anthropoides virgo) portrait, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Four Dwarf irises (Iris pumila) in flower, Rostovsky Nature Reserve Rostov Region, Russia, April 2009
Steppe eagle (Aquila nipalensis) on nest, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Four rare yellow Bieberstein tulips (Tulipa biebersteiniana) in flower, Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Dwarf irises (Iris pumila) flowering, Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Feather grass (Stipa sp) at sunset in the steppe of Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, April 2009
Close-up inside Wild tulip (Tulipa schrenkii) Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
A rare yellow Bieberstein tulip (Tulipa biebersteiniana) Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Wild tulips (Tulipa schrenkii) in flower, one white, the rest red, Rostovsky Nature Reserve, Rostov Region, Russia, April 2009
Pygmy ground squirrel / souslik (Spermophilus pygmaeus) standing on hind legs alert, Cherniye Zemli (Black Earth) Nature Reserve, Kalmykia, Russia, May 2009
Male Red footed falcon (Falco vespertinus) hunting over burning steppe fields, Bagerova Steppe, Kerch Peninsula, Crimea, Ukraine, July 2009 WWE OUTDOOR EXHIBITION