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Duckling swimming on water surface, captive, UK
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
A split level digital composite showing a Basking shark (Ceterhinus maximus) feeding on plankton around St Michaels Mount, Cornwall, UK. June
Duckling swimming on water surface, split level, captive, UK
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Florida manatee (Trichechus manatus latirostris) swimming into a fresh water spring, Three Sisters Spring, Crystal River, Florida, USA
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
Split level photo of an American crocodile (Crocodylus acutus) beneath red mangrove trees (Rhizophora mangle) above a bed of seagrass (turtlegrass: Thalassia testudinum)
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Split level view of blue green waters of Verzasca River flowing over granite rocks at Lavertezzo, Canton Tessin, Switzerland, September. Taken for the Freshwater Project
Grey seal (Halichoerus grypus) split level of young seal investigating the camera with its nose and whiskers, close to shore. Lundy Island, Devon, UK, Bristol Channel. August
Pair of Sockeye salmon (Oncorhynchus nerka) on their redd in a shallow stream. Female in front of male. Huihil Creek, Adams River, British Colombia, Canada, October
A split level image of Southern stingray (Hypanus americanus) swimming over a sand bar. Grand Cayman, Cayman Islands. Caribbean Sea. Digital composite
Juvenile Green turtle (Chelonia mydas) swimming near the surface, split level view, Fakarava atoll lagoon, Tuamotu Archipelago, French Polynesia, Pacific Ocean
White waterlily (Nymphaea alba) in Naardermeer bog lake, Holland. May
Three Sockeye salmon/ Red salmon (Oncorhynchus nerka) swimming upstream as they migrate back to the river of their birth to spawn
A sprouting coconut floating in the sea close to the shore, Yap, Micronesia, Pacific Ocean
RF - Split level photo of mangrove scenery, with hard corals (including Goniopora sp. Porites sp.) growing below Red mangrove tree (Rhizophora sp.). Raja Ampat, West Papua, Indonesia
Southern stingray (Dasyatis americana) two swimming over sand bar, under stormy sky. Grand Cayman, Cayman Islands. British West Indies
Chum salmon (Oncorhynchus keta) migrate up a small river near Bella Bella, British Columbia, Canada. September
Guillemot (Uria aalge) swimming, split level view, Shiant Isles, Outer Hebrides, Scotland, UK. June, 2018
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Split level view of water in cave in Wadi Al Shab, Al Sharqiyah South Governorate, Sultanate of Oman. February. Photographed for The Freshwater Project
White waterlily (Nymphaea alba) in a peat bog lake, De Weerribben-Wieden National Park, Holland. September
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF - Split level photo of a Whale shark (Rhincodon typus) feeding at the surface. Isla Mujeres, Yucatan Mexico. Caribbean Sea
Southern stingrays (Hypanus americanus) swimming over a sand bar in the early morning. Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Southern stingray (Hypanus americanus) at the surface. Stingray City, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea. Bait was used for this shot
Florida manatee (Trichechus manatus latirostrus) with Blue gill sunfish (Lepomis macrochirus) cleaning it, in a freshwater spring, beneath trees. Three Sisters Spring, Crystal River, Florida, USA
Split level photo of an American crocodile (Crocodylus acutus) floating at the surface over a shallow seagrass meadow, close to mangroves
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Lemon shark (Negaprion brevirostris) in shallow water at the surface, split level. Little Bahama Bank, Bahamas. West Atlantic Ocean
Split level image of Caribbean reef shark (Carcharhinus perezi) breaking surface at dusk. Grand Bahama, Bahamas. Digitally manipulated image
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Spectacled caiman (Caiman crocodilus) Rio BaiAia Bonita, Bonito, Mato Grosso do SulSpectacled caiman (Caiman crocodilus) Rio Baiia Bonita, Bonito, Mato Grosso do Sul, Brazil
Dalmatian pelicans (Pelecanus crispus) low angle perspective of open bills, Lake Kerkini, Greece, February. Highly Commended, 2011 Wildlife Photographer of the Year competition
Couple of Pink River dolphin / Boto (Inia geoffrensis) Acajatuba Lake, Negro River, Amazonas, Brazil
Coral reef with Threadfin anthias (Pseudanthias huchtii) shoal with Bluegreen chromis (Chromis viridis) amongst them, under shallow sea with a coconut tree forest in background, Philippines
Two Sockeye / Red Salmon (Oncorhynchus nerka), female digging riverbed to lay eggs on spawning ground. Trees showing autumnal colours, Adams river, British Columbia, Canada. October
Split-level of a Sperm whale (Physeter macrocephalus) swimming close to the surface, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit. February
Two Florida manatees (Trichechus manatus latirostris) close to the surface in shallow water, Three Sisters Spring, Crystal River, Florida, USA. Endangered
Coral reef below the cliffs of Shark Observatory, Ras Mohammed National Park, Sinai, Egypt, Red Sea
Portuguese man o war (Physalia physalis) floating on sea surface. Azores, Atlantic Ocean