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Lemon (Citrus limon) flower, buds and fruit, Umbria, Italy. June
Apennine treacle mustard (Erysimum pseudorhaeticum) and Greater milkwort (Polygala major) in flower in dry field, Gran Sasso, Abruzzo, Italy. June
Fallow fields burst into flower containing Corn chamomile (Anthemis arvensis), Hoarry mullein (Verbascum pulverulentum), Purple toadflax (Linaria purpurea) and Common poppy (Papaver rhoeas), Abruzzo
Fallow field in flower containing Poppy (Papaver rhoeas), Woad (Isatis tinctoria), Corn chamomile (Anthemis arvensis) Olympian mullein (Verbascum longifolium) and Purple toadflax (Linaria purpurea)
Santa Fe land iguana (Conolophus pallidus) with Giant prickly-pear cactus (Opuntia echios), Santa Fe Island, Galapagos, South America
Galapagos carpetweed (Sesuvium edmonstonei) and giant Opuntia (Opuntia sp)Galapagos carpetweed (Sesuvium edmonstonei) and giant Opuntia (Opuntia sp.) cacti, beneath a rainbow at sunset, Plazas Island, Galapagos, South America
Galapagos land iguana (Conolophus subcristatus) recently reintroduced to Santiago island where it had become extinct, feeding on Yellow cordia (Cordia lutea) leaves, Santiago Island, Galapagos
Darwins medium ground finch (Geospiza fortis) feeding on wild seeds (Tournefortia sp), Santa Cruz Island, Galapagos, South America
Darwins large ground finch (Geospiza magnirostris) feeding on Wild cucumber (Momordica sp), Santa Cruz Island, Galapagos, South America
Koala (Phascolarctos cinereus) eating gum (Eucalyptus sp)Koala (Phascolarctos cinereus) eating gum (Eucalyptus sp.) leaves in a tree, Apollo Bay, Victoria, Australia
Eucalypt rainforest along Otway Lighthouse Road, Cape Otway, The Otways, Victoria, Australia. April 2021
House finch (Carpodacus mexicanus) perched on Saguaro cactus (Carnegiea gigantea) in flower, Sonoran Desert, Arizona
Female European hoverfly (Helophilus pendulus) pollinating Tansy (Tanacetum vulgare) in flower, Belgium. August
Lesser paper wasp (Polistes dorsalis) on Goldenrod (Solidago sp)Lesser paper wasp (Polistes dorsalis) on Goldenrod (Solidago sp.) flower, Philadelphia, Pennsylvania, USA
Gila woodpecker (Melanerpes uropygialis) feeding on Saguaro (Carnegiea gigantea) blossom nectar and the insects trapped in them, Sonoran Desert, Arizona, USA
A large herbivorous Seahare (Aplysia punctata) crawling around a Common eelgrass (Zostera marina) meadow, Helford River Estuary, Cornwall, UK
Carrion fly on the parasitic plant Rafflesia pricei flower, Mount Kinabalu, Borneo, May, 2013. Filmed for the BBC series Wonders of The Monsoon
Four Hereford bullocks looking over a fence with Cow Parsley (Anthriscus sylvestris) Worcestershire, England, May
White stork (Ciconia ciconia) flying back to its nest in Oak (Quercus robur) tree to feed its three large chicks, Knepp Estate, Sussex, UK, June
Male Common Redpoll (Acanthis flammea) perched on snow-covered dead stem of Evening Primrose (Oenothera biennis), from which it will eat seeds, New York, USA
Male Pine grosbeak (Pinicola enucleator) perched on branch among Crabapple (Malus sp) fruits, near Scenectady, New York, USA
Flower-rich grassland with Red clover (Trifolium pratense) beside Jurassic Coast path, near Swanage, Dorset, UK, July 2021
Red giant tajinaste / Mount Teide bugloss / Towers of jewels (Echium wildpretii) close up of flowers, Teide National Park, Tenerife, Canary Islands
Gilded flicker (Colaptes chrysoides) flying to feed on nectar from Saguaro blossoms (Carnegiea gigantea), Sonoran desert, Arizona
Bumblebee (probably Bombus terrestris) visiting flowers of Sainfoin (Onobrychis viciifolia) in a restored wildflower meadow, near Bristol, UK, June
Ghost orchid (Epipogium aphyllum) flowering in leaf litter on forest floor, Roncal Valley, Navarre, Spain, Europe
Ghost orchid (Epipogium aphyllum), Roncal Valley, Navarre, Spain, Europe
Eastern spinebill (Acanthorhynchus tenuirostris) male visiting Grevillea flower, small honeyeater found in forests, woodlands and gardens in Eastern Australia, Toowoomba, Queensland, Australia
Regent bowerbird (Sericulus chrysocephalus) male sitting in budding mango tree (Mangifera indica), Ravensbourne, Queensland, Australia
Downy Birch (Betula pubescens) showing autumn colours. Glen Strathfarrar, Scottish Highlands. Scotland. October
Beech (Fagus sylvatica), large tree amongst Bracken (Pteridium aquilinum). Bolderwood, New Forest National Park, Hampshire, England, UK. October 2020
Autumn leaves, Japanese maple (Acer palmatum) Lu Shan, Taiwan
Autumn colours in Mountain birch, (Betula pubescens var. tortuosa), Abisko National Park, Norrbotten, Lapland, Sweden September 2020
Autumn leaves including Sweet gum (Liquidambar styraciflua); Hawthorn (Crataegus monogyna); Tutsan (Hypericum androsaemum); Norway maple (Acer platanoides)
Aspen (Populus tremula) Wester Ross, Scotland, UK. November
Raindrops on fallen autumn quaking aspen leaves (Populus tremuloides), Colorado, USA. October
European beech forest (Fagus sylvatica) in autumn, view from below in the mist, Ilirska Bistrica, Green Karst, Slovenia, October
Roe deer (Capreolus capreolus) feeding in field with flowering Poppies (Papaver rhoeas) and Cornflower (Centaurea cyanus), De Inslag, Brasschaat, Belgium, June
Aguaje palm swamp (Mauritia flexuosa) Maijuna Indigenous Community, Rainforest, Sucusari, Rio Napo, Loreto, Peru. January 2013
Oak tree (Quercus robur) growing among bracken (Pteridium aquilinum), autumn colour, The New Forest, Hampshire, UK. November 2016
Beech trees (Fagus sylvatica) and English bluebells (Hyacinthoides non-scripta) soft focus, West Wood, near Marlborough, Wiltshire, UK. May 2021
RF - Beech trees (Fagus sylvatica) and english bluebells (Hyacinthoides non-scripta). Late evening light and double exposure to create soft, dreamy effect. West Woods, nr Marlborough, Wiltshire, UK
Luskentyre beach/sands, marram grasses and early morning sunlight, Isle of Harris, Outer Hebrides, Scotland, UK. October 2018
Common spotted orchids (Dactylorhiza fuchsii), Kingcombe Meadows, Dorset, UK, June
Cow parsley (Anthriscus sylvestris) in a misty Crendle Woods at dawn, Milborne Port, Somerset, England, UK. May 2020
Gila woodpecker (Melanerpes uropygialis) pair at nest entrance in Saguaro cactus (Carnegiea gigantea) one adult flying from nest, Sonoran Desert, Arizona, USA
Gilded flicker (Colaptes chrysoides) feeding on Saguaro cactus (Carnegiea gigantea) blossom nectar, Sonoran desert, Arizona, USA
Altai pansy (Viola altaica) Valley of Petrified Clouds. Altai Republic, Russia