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Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Eastern grey kangaroo (Macropus giganteus), two males fighting. Grampians National Park, Victoria, Australia
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK