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Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Young ground squirrels (Xerus inauris) playing, Kgalagadi Transfrontier Park, South Africa. Non-ex
Welsh Ponies (Equus caballus) engaged in mutual grooming, Llanbedr Hill, Powys, Wales, August
Impala (Aepyceros melampus) allo-grooming, iMfolozi game reserve, KwaZulu-Natal, South Africa, August
Pack of wolves (Canis lupus). Yellowstone National Park, Wyoming, USA. January
Lions (Panthera leo) five drinking together, Ndutu, Tanzania
Lionesses (Panthera leo) grooming each other, Masai-Mara Game Reserve, Kenya
Two white horses of the Camargue, interacting, greeting head to head in marshes, Camargue, Southern France
Common / Eurasian crane (Grus grus) flock in flight with snow topped Pyrenees mountains in the background, during southward migration period, Haute Pyrenees, Gascony, France. December 2009
African lions (Panthera leo) one lioness licking another, Okavango Delta, Botswana
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Wild Chincoteague (Equus caballus) two breeding stallions greeting one another, Chincoteague National Wildlife Refuge, Chincoteague Island, Virginia, USA, June
Black-tailed Prairie Dogs (Cynomys ludovicianus), two pups (right) seem to be kissing their parent. The behaviour is a gesture of recognition
Sperm whales (Physeter macrocephalus) large group gathering together, rubbing together to shed skin, interacting and defecating. Indian Ocean, March
Aggregation of Sperm whales (Physeter macrocephalus) several females swim after a few males, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit. February
Long-eared owls (Asio otus) autumn, two owls interacting while roosting in tree, social behaviour, The Netherlands
Pair of European bee-eaters (Merops apiaster) with courtship offering of insect prey, Pusztaszer, Kiskunsagi National Park, Hungary
White-faced Capuchins (Cebus capucinus imitator) grooming while resting.Osa Peninsula, Costa Rica
Formosan rock macaque (Macaca cyclopis) social grooming behaviour. Alishan National Recreational Forest, Taiwan
Formosan macaques (Macaca cyclopis) social grooming, Taiwan. Endemic
Two wild Konik horse stallions greeting one another, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Southern right whales (Eubalaena australis) aerial view of three adullts engaged in social activity, photographed with the permission of the Department of Environmental Affairs, South Africa
Group of Sperm whales (Physter macrocephalus) family group swimming together, Indian Ocean, March
Aggregation of Sperm whales (Physeter macrocephalus) engaged in social activity. Indian Ocean, March
Large aggregation of Sperm whales (Physeter macrocephalus) engaged in social activity. Note the abundance of sloughed off skin in the water, resulting from tactile contact among the whales
Ruffs displaying (Philomachus pugnax) Agapa River, Taimyr Peninsula, Siberia, Russia
Cheetah (Acinonyx jubatus) two leaning close together, Okavango Delta, Botswana
Black-Mantled Tamarin (Saguinus nigricollis) group with social grooming, Sumaco, Napo, Ecuador
Honeybee workers exchanging food - known as trophallaxis (Apis mellifera) Sussex, UK
Flock of Cattle Egrets (Bubulcus ibis) in roosting trees at dusk, Imba Ndutu, Serengeti NP, Tanzania, East Africa
Large group of Californian sealions (Zalophus californianus) swimming underwater, Anacapa Island, California, Pacific Ocean
Small flock of Greater flamingo (Phoenicopterus roseus) wading on inland Kalloni lake, Lesbos, Greece
Bat rays (Myliobatis californica) schooling, Redondo Rock, Galapagos Is
Leaf cutting ants (Atta cephalotes) cutting pieces of leaf, rainforest habitat, Costa Rica
