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Male Black grouse (Tetrao tetrix) displaying at lek at dawn, Cairngorms NP, Grampian, Scotland, UK, April
Short-eared owl (Asio flammeus) in flight, backlit, at dusk, Lincolnshire, UK, March
European hare (Lepus europaeus) running across game cover on edge of large arable field at dawn, Norfolk, England, UK, April
Curlew (Numenius arquata) in flight against sky at dusk, Peak District NP, UK, March 2011
Two Common Coot (Fulica atra) swimming and leaving wakes on water. The Netherlands, July
Flock of Pink-footed geese (Anser brachyrhynchus) flying on migration at sunset, Martin Mere WWT, Lancashire, UK, September
African skimmer skimming at sunset {Rynchops flavirostris} Chobe NP, Botswana
Adult Violin Beetle (Mormolyce borneensis) on decaying leaf on the rain forest floor. Near Ginseng Camp in the heart of Maliau Basin - Sabahs Lost World. Maliau Basin, Borneo
Silhouette of five leaping Children, Bohuslan archipelago, Sweden
Goliath heron (Ardea goliath) near Mopani Dam at dusk, Kruger National Park, Transvaal, South Africa, September
Masai giraffe (Giraffa camelopardalis) and her youngster on the savanna landscape at dawn, Serengeti National Park, Tanzania
Silhouette of Red-fronted brown lemur (Eulemur rufus) leaping across gap in trees by moonlight, cathemeral behaviour, Ranomafana National Park, south east Madagascar, November
Arctic Tern (Sterna paradisaea) in flight, silhouetted against the moon at dusk, Uto Finland July
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Whitetip reef sharks (Triaenodon obesus) pack silhouetted following scent trail in water column, Cocos Island, Costa Rica, Pacific Ocean
Giraffe (Giraffa camelopardalis) three standing together, silhouetted at dusk, Okavango Delta, Botswana
Red Deer (Cervus elaphus) hinds silhouetted in grassland. Richmond Park, London, UK, October
Alpine Chough / Yellow-billed Chough (Pyrrhocorax graculus) flock in flight against mountains, Dolomites, Italy. Digital composite
Narwhal (Monodon monoceros) crossing tusks above water surface. Baffin Island, Nunavut, Canada, April
Huge flock of Common starlings (Sturnus vulgaris) flying in to roost over the reed beds at dusk, Ham Wall nature reserve, Somerset, UK, January
Common raven (Corvus corax) perched in snow beside birch tree, Finland, March
Killer whale / Orca (Orcinus orca) surfacing with three seabirds flying, Tysfjord, Norway, November
Caribbean reef shark (Carcharhinus perezi) over sand ripples, Walkers Cay, Bahamas. Tropical West Atlantic Ocean. Taken in natural light
Herd of Wildebeest (Connochaetes taurinus) silhouetted at dusk, Serengeti NP, Tanzania, East Africa, January
Crested Newt (Triturus cristatus carnifex) underwater view of female swimming on surface, Traversari lake, Campigna National Park, Emilia Romagna and Tuscany regions, Italy
Silhouette of Moai statue at dawn in Ahu Ko Te Riku, restored archaeological site of Ahu Tahai, Hanga Roa, Easter Island, Pacific ocean, November 2004
Silhouette of Black-winged stilt (Himantopus himantopus) standing on exposed rock over water, with reflections, Lesbos, Greece
Ring-tailed Lemur (Lemur catta) sun basking at dawn. Berenty Private Reserve, southern Madagascar
Moose (Alces alces) bull shaking off water at sunset, Great North Woods, Baxter State Park, Maine, New England, USA, Autumn
Sillhouette of Black woodpecker {Dryocopus martius} flying from nest hole in tree trunk in mist / rain, ancient forest, Vosges mountains, Lorraine, France
Wildebeest (Connochaetes taurinus) running, Tanzania
Grulla Mustang stallion running in dust at Return to Freedom Sanctuary, Lompoc, California, USA
Jabiru stork (Jabiru mycteria) pair on nest at sunset, Pantanal NP, Mato Grosso, Brazil
Red Deer (Cervus elaphus) stag silhouetted in mist, Dyrehaven, Denmark
Looking up at Baobabs (Andasonian grandidieri) on Baobabs Avenue, Morondava, West Madagascar
Silhouette of three cowboys and a dog riding down a ridge, Flitner Ranch, Shell, Wyoming, USA, model released
Bluefin tuna {Thunnus thynnus} in fish farm, Kusimoto, South Japan
Spotted Hyaena (Crocuta crocuta) at night, South Africa
Silhouette of White storks {ciconia ciconia} nesting on purpose-built poles, Malpartida de Caceres, Extremadura, Spain Note - relocated after construction of hotel
Three Mute Swans (Cygnus olor) on frozen loch in early morning mist, Cairngorms National Park, Scotland, UK
Silhouette of Eastern Grey Kangaroo {Macropus giganteus} males boxing, New South Wales, Australia
Lesser black noddies {Anous tenuirostris} nesting, silhouetted, Noronha Is, Brazil
Tourists watching Mount Yasur erupt at dusk, Tanna, Vanuatu Aug-04
Pacific manta ray {Manta alfredi} Yap, Micronesia
Common zebra at sunset. (Equus quagga). Okavango, Botswana
Silhouette of King penguin {Aptenodytes patagonicus} standing tall during display, at sunset, Falkland Islands
English Oak (Quercus robur) silhouetted on horizon at sunrise. West Milton, Dorset, England, UK, February
Pico do Areeiro, Madeira Island, April 2009
