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Tompot blenny (Parablennius gattorugine) in bright summer mating colours, peering out from his home in a discarded tube, where he was guarding his eggs (not visible in this photo), Swanage Pier
St Abbs harbour (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, August 2011
Bottlenose dolphin (Tursiops truncatus) two breaching in evening light, Moray Firth, Inverness-shire, Scotland, UK, August, sequence 2/2
Portrait of Ballan wrasse (Labrus bergylta), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October 2011
Bass Rock at dawn, North Berwick, Scotland, UK, August. 2020VISION Book Plate
Atlantic Puffin (Fratercula artica) adult on breeding cliffs. Hermaness NNR, Shetland, UK, June
Puffin (Fratercula arctica) swimming underwater, Farne Islands, Northumberland, UK, July
Arctic tern (Sterna paradisaea) in flight, Inner Farne, Farne Islands, Northumberland, June
Gannet (Morus bassanus) portrait, Bass Rock, Firth of Forth, Scotland, UK, June
Flock of Manx shearwaters (Puffinus puffinus) in flight over sea, near Ardnamurchan Point, Inner Hebrides, Scotland, UK, July 2011
Patterns in sand, North Harris, Outer Hebrides, Scotland, UK, October 2010
Amphipod (Iphimedia obesa) on top of a colony of Deadmans fingers (Alcyonium digitatum) coral, Loch Carron, Ross and Cromarty, Scotland, UK, April
Catch of Atlantic mackerel (Scomber scombrus) in fish separator on board Shetland pelagic trawler Charisma, Shetland Isles, Scotland, UK, October 2012
Fraserburgh pelagic trawler Forever Grateful fishing for Atlantic mackerel (Scomber scombrus) close to Papa Stour, Shetland Isles, Scotland, UK, October 2012
Male Black faced blenny (Tripterygion delaisi), part of a sequence showing colour change, Swanage, Dorset, England, UK, May. 2/4
Shanny (Lipophrys pholis) in a crevice in a harbour wall, Bovisand Harbour, Plymouth, Devon, England, UK, July
John dory (Zeus faber), Porthkerris Cove, Cornwall, England, UK, June
Grey seal (Halichoerus grypus) in a gully beneath kelp (Laminaria digitata), Farne Islands, Northumberland, England, UK, July
Basking shark (Cetorhinus maximus) feeding at the surface on plankton, Cairns of Coll, Inner Hebrides, Scotland, UK, June
Arctic tern (Sterna paradisaea) in flight at twilight using slow shutter speed to accentuate blur and motion in low-key light, Inner Farne, Farne Islands, Northumberland, July
Arctic Tern (Sterna paradisaea) with sand eel in beak. Farne Islands, Northumberland, July
Thong seaweed (Himanthalia elongata) and various other seaweeds in shallow water, Cairns of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, August
Kelp forest (Laminaria sp) growing beneath the cliffs of Lundy Island, Devon, UK, Bristol Channel, July
Tompot blenny (Parablennius gattorugine) beneath Swanage Pier, Dorset, UK. Photographed during May. English Channel
Common dolphins (Delphinus delphis) bow-riding, near South Uist, Outer Hebrides, Scotland, UK, June
Common dolphin (Delphinus delphis) at surface, near South Uist, Outer Hebrides, Scotland, UK, June
Adult Kittiwake (Rissa tridactyla) in winter plumage in flight alongside the pelagic trawler Charisma, Shetland Isles, Scotland, UK, October
Atlantic grey seal (Halichoerus grypus) in Lerwick harbour, Shetland Islands, Scotland, UK, October
Northern gannet (Sula bassana) being mobbed by immature Great black-backed gull (Larus marinus), Shetland Islands, Scotland, UK, October
Two Killer whales (Orcinus orca) with a solitary Fulmar (Fulmarus glacialis) seen from the pelagic trawler Charisma in evening light, Shetland Islands, Scotland, UK, October 2011
Atlantic wolf fish (Anarhichas lupus) in a rock crevice, St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, August 2011
Edible crab (Cancer pagurus), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, August 2011
Common Sunstar (Crossaster papposus), St Abbs (St Abbs and Eyemouth Voluntary Marine Reserve), Berwickshire, Scotland, UK, October 2011
Adult Atlantic puffin (Fratercula arctica) in flight in summer, backlit, Isle of Lunga, Inner Hebrides, Scotland, UK, July 2011. 2020VISION Book Plate
RF- Rear view of Basking shark (Cetorhinus maximus) feeding on plankton, visible as white dots on water surface near island of Coll, Inner Hebrides, Scotland, UK. June
Phosphorescent Seapen (Pennatula phosphorea) extends out of the muddy seabed in the calm waters of the upper reaches of a Scottish sea loch. Loch Duich, Ross and Cromarty, Scotland, UK, April
Two European river otters (Lutra lutra) swimming in shallow water, Isle of Mull, Inner Hebrides, Scotland, UK, December. Did you know? 12% of breeding otters in the UK live in the Shetland Isles
European river otter (Lutra lutra) eating fish, resting on seaweed, Isle of Mull, Inner Hebrides, Scotland, UK, December
Basking shark (Cetorhinus maximus) feeding on plankton in the surface waters around the island of Coll, Inner Hebrides, Scotland, UK, June. 2020VISION Book Plate
Puffin (Fratercula arctica) swimming underwater, Farne Islands, Northumberland, UK, North Sea, July
Mollusc shells washed up on a beach in the Cairns of Coll, Island of Coll, Inner Hebrides, Scotland, UK, North Atlantic Ocean, June 2011
Detail of Plumose anemone (Metridium senile) in its retracted state at slack water when it is not feeding. Plymouth Sound, Devon, England, UK, English Channel, July
Devonshire cup coral (Caryophyllia smithii) photographed under deep blue light to reveal its fluorescent pigments, showing up as green in this photo. Plymouth Sound, Devon, UK, English Channel, July
Polar bear (Ursus maritimus) female and her two cubs on large iceberg off Baffin Island. Nunavut, Canada, August 2010
Detail of the filter feeding apendange of Peacock worm (Sabella pavonina) Loch Carron, Ross and Cromarty, Scotland, UK, April
Bottlenose dolphin (Tursiops truncatus) group surfacing in evening light, Moray Firth, Inverness-shire, Scotland, UK, September
Grey seal (Halichoerus grypus) portrait, underwater amongst kelp, Farne Islands, Northumberland, England, UK, August
Common Lobster (Homarus gammarus), in a rock crevice, Lundy Island Marine Conservation Zone, Devon, England, UK, May. Did you know