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Reindeer (Rangifer tarandus) female near Aviemore, Cairngorms National Park, Scotland, UK, December
Traigh Scarasta on the Isle of Harris. Isle of Harris, Outer Hebrides, Scotland, UK, August
Clashach cove surrounded by flowering Gorse (Ulex europaeus) Hopeman, Scotland, UK, May
Crested tit (Lophophanes cristatus) perched above pine forest with a series of small lochans in background, Cairngorms National Park, Scotland, UK, December 2014
Impression of Birch tree (Betula pendula) in autumn, Scotland, UK. November
Silver birch (Betula pendula) trees in early autumn, Craigellachie National Nature Reserve, Aviemore, Cairngorms National Park, Scotland, UK, September
Eurasian red squirrel (Sciurus vulgaris) reflected in water, Cairngorms National Park, Scotland, United Kingdom, October
Red squirrel (Sciurus vulgaris) foraging in forest, climbing down tree trunk, Cairngorms National Park, Scotland, October
European larch (Larix decidua) trees in mist, Scotland, UK, October
Silver birch (Betula pendula) woodland in autumn, Glen Moriston, Scotland, UK, October 2011
Silver birch (Betula pendula) leaves in autumn, Glen Affric, Highland, Scotland, UK, October
Rainbow over Loch Beinn a Mheadhoin, Glen Affric, Highland, Scotland, UK, October 2010
Scattered Scots pine trees (Pinus sylvestris) beside the Dorback Burn, Dorback Estate, Cairngorms National Park, Scotland, UK, September 2011
Grey squirrel (Sciurus carolinensis) foraging in an oak woodland. Perthshire, Scotland, Nov
Silver birch trees {Betula pendula} in autumn, Glen Strathfarrar NNR, Scotland, UK, October
Wild cat (Felis sylvestris) portrait, part of captive breeding project at Alladale Estate, Scotland, UK, October
Sequence of strips of cultivated Machair with early shoots of Black oats (Avena strigosa) pushing through the sand. North Uist, Scotland, UK, June
Corncrake (Crex crex) walking at dawn on Machair, North Uist, Scotland, UK, June
Two Little terns (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Little tern (Sterna albifrons) at the nest amongst Black oats (Avena strigosa) growing on shell rich sands Machair, North Uist, Scotland, UK, June
Corn Marigold (Chrysanthemum segetum) and Scentless mayweed (Tripleurospermum inodorum) machair habitat, North Uist, Scotland, UK, July
Red Deer (Cervus elaphus) portrait of head of stag and antlers. Alladale Estate, Scotland, UK, October
Partially wooded uplands with Five Sisters of Kintail in background, Northwest Highlands, Scotland, UK.September
Purple loosestrife (Lythrum salicaria) flower spikes in evening light, Scotland, UK, September
Hermit crab (Pagurus bernhardus) on yellow sponge, Loch Etive, west coast of Scotland, UK, September
Common Toad (Bufo bufo) juvenile toadlet walking over moss in birchwood in late summer, Inverness-shire, Scotland, September
Eurasian otter (Lutra lutra) feeding whilst on log in woodland burn, Glenfeshie, Cairngorms National Park, Scotland, UK, September
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Rushes reflected in water at dawn, Scotland, UK, September
Red deer (Cervus elaphus) male stag silhouetted at dusk, rut season, Jura, Scotland, UK, September
Rock archway at sunset, Isle of Lewis, Outer Hebrides, Scotland, UK, September 2014
Horse mussel (Modiolus modiolus) bed with Brittlestars (Ophiothrix fragilis) and Edible crab (Cancer pagurus), Shetland, Scotland, UK, September
Loch Garten and Abernethy Forest. Cairngorms National Park, Scotland, UK, September 2011
Barn Swallow (Hirundo rustica) feeding a fledgling on a wire. Perthshire, Scotland, September
Bottlenose dolphin (Tursiops truncatus) group surfacing in evening light, Moray Firth, Inverness-shire, Scotland, UK, September
Pine Marten (Martes martes) standing alert on a trunk. Cairngorms National Park, Scotland, September
Sparrowhawk (Accipiter nisus) watching for prey from a garden rowan tree. Perthshire, Scotland, September
Footpath leading up to Ciore Lagan and Sgurr Sgumain, above Glenbrittle. Cuillin Hills / MtsFootpath leading up to Ciore Lagan and Sgurr Sgumain, above Glenbrittle. Cuillin Hills/Mts, Isle of Skye, Scotland. September 2010
Rannoch Moor on a rainy day. Highlands, Scotland. September 2010
Dew on cobweb, Berwickshire, Scotland, UK, September
RF - Black Grouse (Tetrao tetrix) male peforming flutter jump display on lek, Cairngorms National Park, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
RF - Brown Hare (Lepus europaeus) running through field of grass, Scotland, UK.May (This image may be licensed either as rights managed or royalty free.)
Red deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stags silhouetted at sunset in winter, Scotland, UK.February
Sunset over the Cullin Mountains from Elgol shoreline, Isle of Skye, Scotland, UK, April
Mountain hare footprints in snow on mountain top in Glen Coe, Lochaber, Scotland, UK.February
Crested tit (Parus cristatus) backlit on lichen-covered branch, Inshriach Forest, Scotland, UK, January.January
River Spey on winters dawn, Cairngorms National Park, Scotland, UK. JanuaryRiver Spey on winters dawn, Cairngorms National Park, Scotland, UK.January
Buacaille Etive Mor and River Etive in winter, Rannoch Moor, Glencoe, Scotland, UK.December