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School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands
Spotted eagle ray (Aetobatus narinari) Kicker Rock, San Cristobal Island, Galapagos
Galapagos penguins (Spheniscus mendiculus) underwater with shoal of fish, Tagus Cove, Isabela Island, Galapagos
Scalloped hammerhead shark shoal (Sphyrna lewini) Wolf Island, Galapagos
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Striped marlin (Kajikia audax) hunting Sardine (Sardinops sagax) in bait ball. Magdalena Bay, Baja California Sur, Mexico
School of Cow breams (Sarpa salpa ) Azores, Atlantic ocean
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Munks pygmy devil ray / Munks mobula (Mobula munkiana) large school from the air, Baja California, Mexico. Large group gathered possibly for mating purposes
Blacksmiths (Chromis punctipinnis) school swam through a giant kelp (Macrocystis pyrifera) forest, Santa Barbara Island, Channel Islands. Los Angeles, California, USA, North East Pacific Ocean
Nurse shark (Ginglymostoma cirratum) and Horse-eye jacks (Caranx latus) Shark Ray Alley, Hol Chan Marine Reserve, Belize Barrier Reef, Belize
Gorgonian / Sea whip coral (Ellisella ceratophyta) with Ring tailed cardinalfish (Apogon aureus) swimming in front, West Papua, Indonesia
Coral reef with soft corals (Dendronephthya sp. ), encrusting sponges and Pygmy sweepersCoral reef with soft corals (Dendronephthya sp.), encrusting sponges and Pygmy sweepers (Parapriacanthus ransonetti) Similan Islands, Andaman Sea, Thailand
School of Perch (Perca fluviatilis) amongst Water lilies (Nuphar lutea) lit by the suns rays, Lake Bourget, Alps, Savoie, France, June
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Cichlids (Cichlidae) in Lake Malawi, Malawi, November. Photographed for The Freshwater Project
Striped mackerel (Rastrelliger kanagurta) shoal swimming with open mouths, filtering for zooplankton. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Snorkeller with shoal of Piraputanga (Brycon hilarii), Rio Olha d Agua at Cabeceira do Prata, Bonito, Mato Grosso do Sul, Brazil
Oceanic whitetip shark (Carcharhinus longimanus) accompanied by a group of Pilotfish (Naucrates ductor). Rocky Island, Egypt. Red Sea
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
The vertical reef wall at Shark Reef, Ras Mohammed, with Scalefin anthias (Pseudanthias squamipinnis) and soft corals (Dendronephthya spp.) Yellowsaddle goatfish (Parupeneus cyclostomus)
School of Longnose parrotfish (Hipposcarus harid) swarming over reef. Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
RF- School of Blackfin barracuda (Sphyraena qenie) forming circle in open water at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
RF - A vibrant Red Sea coral reef scene, with orange female Scalefin anthias fish (Pseudanthias squamipinnis) teeming over Fire coral (Millepora dichotoma)
Diver swimming with an aggregation of Cod (Gadus morhua) over a kelp forest. These cod were gathered in early spring off the north coast of Iceland to spawn. Thorshofn, Iceland. North Atlantic Ocean
Blue and gold fusilier / Scissor tailed fusilier (Caesio caerulaurea) school swimming through shafts of sunlight, West Papua, Indonesia
RF - School of large Atlantic bluefin tuna (Thunnus thynnus) captive in a growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Blue spotted trevally (Caranx bucculentus) hunting small fish in the reef, South Ari Atoll, Maldives, Indian Ocean
School of powderblue surgeonfish (Acanthurus leucosternon) feeding, Laamu Atoll, Maldives. Indian Ocean
Slate pencil sea urchins (Heterocentrotus mammillatus) on coral reef with Pyramid butterflyfish (Hemitaurichthys polylepis), Hawaii
Loggerhead turtle (Caretta caretta) accompanied by pilotfish (Naucrates ductor) swimming near the surface. Santa Maria Island, Azores, Portugal, Atlantic Ocean. Vulnerable
School of Oriental sweetlips (Plectorhinchus vittatus) gather tightly together as they rest during the day, with a Clown triggerfish (Balistoides conspicillum)
