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Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Fosa (Cryptoprocta ferox) male marking territory onto a mating tree, Kirindy Forest, Madagascar
Asian black bear (Ursus thibetanus) scent marking tree by rubbing back on it, Land of the Leopard National Park, Russian Far East. Taken with remote camera. December
Siberian tiger (Panthera tigris altaica) sitting and scent marking tree in forest by rubbing body against it, Land of the Leopard National Park, Russian Far East. Endangered
Lion (Panthera leo) male, scent marking its territory at dawn, Ngorongoro Conservation Area / Serengeti National Park boundary, northern Tanzania
Iberian lynx (Lynx pardinus) sniffing scent on dead branch, Parque Natural Sierra de Andujar, Andalucia, Spain. January