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Chocolate chip sea star o (Protoreaster nodosus) on sandy bottom. Mabul, Malaysia
Tube anemone (Cerianthus sp) Lembeh, Sulawesi, Indonesia
Emperor angelfish (Pomacanthus imperator) swimming past coral reef. Egypt, Red Sea
Yellowbar angelfish (Pomacanthus maculosus) Egypt, Red Sea
Gorgonian sea fan (Melithaea sp. ) with Lyretail anthias (Pseudanthias sqamipinnis)Gorgonian sea fan (Melithaea sp.) with Lyretail anthias (Pseudanthias sqamipinnis). Similan Islands, Andaman Sea, Thailand
Kaleidoscopic image of Bubble tip anemone (Entacmaea quadricolor). Raja Ampat, West Papua, Indonesia
Kaleidoscopic image of Feather stars (Crinoidea) and Gorgonian wrapper anemone.(Nemanthus annamensis). Indonesia. Western Pacific
Antarctic fur seal (Arctocephalus gazella), mother warning off curious juvenile. Southern elephant seal (Mirounga leonina) in background. Gold Harbour, South Georgia. October
Razorbill (Alca torda) perched on lichen covered rock. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Kittiwake (Rissa tridactyla) flying over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Herring gull (Larus argentatus) flying over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Coumeenoole Bay Slea Head with a view towards Dunmore Head, Dingle Peninsula, County Kerry, Republic of Ireland. November 2013
The Cobb, Lyme Regis, Jurassic Coast, Dorset, England, UK. December 2010
Sand dunes at Luskentyre with view to Taransay. Isle of Harris, Outer Hebrides, Scotland, UK. November 2016
Northern gannet (Morus bassanus), Feeding frenzy, Noss NNR, Shetland, Scotland, UK
Gaada Stack, Foula, Shetland, Scotland, UK. August, 2016
A Kamchatka Brown Bear (Ursus arctos beringianus) standing in breaking waves. Pacific coast of Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
Goldring tunicate (Clavelina robusta) Puerto Galera, Philippines
Polyps on gorgonian fan coral. West Papua, Indonesia
Blue bat star (Patiria pectinifera). Bohai Sea, Yellow Sea. Penglai / Dengzhou / Tengchow, Yantai, Shandong Province, China
Allied cowrie (Diminovula punctata) on a soft coral (Dendronepthya sp ) with its egg mass close by. Po Bin Chau, Sai kung, Hong Kong, China
Gold-spotted Chromodorid nudibranch (Chromodoris aureopurpurea, Ninepin Group or Kwo Chau Islands)Gold-spotted Chromodorid nudibranch (Chromodoris aureopurpurea, Ninepin Group or Kwo Chau Islands, Sai Kung District of Hong Kong, China
Indo-Pacific humpback dolphins (Sousa chinensis) surfacing, Tai O, Lantau Island, Hong Kong, China
Nudibranch (Kaloplocamus acutus) Bluff Island, Hong Kong UNESCO Global Geopark, Hong Kong, China
Sea apple, eating, using arm to transfer food into mouth. Ninepin Group or Kwo Chau Islands, Sai Kung District of Hong Kong, China
Crown spotty blenny (Laiphognathus longispinis) Pak Lap Tsai, Sai Kung East Country Park, Hong Kong, China
Bivalves on the rocky shore, Zhifu Island. Shandong Province, China
Sun / Tube coral (Tubastraea sp) with tentacles extended to catch zooplankton. Gulf of Bohai, Yellow Sea. Penglai / Dengzhou / Tengchow, Yantai, Shandong Province, China
Rocks along Yellow Sea coast at sunset. Yangma Island, Yantai, Shandong Province, China. September 2017
Ascidian / Sea squirt (Ascidiacea) close-up. Bohai Sea, Yellow Sea. Zhifu Island, Shandong Province, China
Blue shark (Prionace glauca) with Pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
American / Northern lobster (Homarus americanus) partially hidden under seaweed, Nova Scotia, Canada, July
Aerial view of the Eastern shore of Nova Scotia, Canada, September
North Atlantic Right Whale (Eubalaena glacialis) tail fluke, in the Bay of Fundy, Canada, September
Minke whale ( Balaenoptera acutorostrata) breaching, Bay of Fundy, Canada, July
Long beaked common dolphin (Delphinus capensis) pod porpoising Baja California, Mexico
Californian Sealion (Zalophus californianus) in kelp (Macrocystis pyrifera) forest, San Benito del Este Island, Baja California Peninsula Pacific Islands Biosphere Reserve, Baja California, Mexico
Split level view of Brown pelicans (Pelecanus occidentalis) feeding, Tortuga Bay, Santa Cruz Island, Galapagos
Green sea turtles (Chelonia mydas) on corals, Hawaii
Andean flamingo (Phoenicoparrus andinus) feeding on shore Laguna Hedionda, Altiplano, Bolivia
Jamess flamingo (Phoenicoparrus jamesi) flock on the shore of Laguna Colorada / Reserva Eduardo Avaroa, Altiplano, Bolivia
Sargassum fish (Histrio histrio) at the sea surface with floating sargassum weed. Hawaii
Painted cardinalfish (Archamia fucata) Yap, Micronesia
European barracuda (Sphyraena sphyraena) and Bluefish (Pomatomus saltatrix) circling baitball of Atlantic horse mackerel (Trachurus trachurus) Formigas Islets, Azores
Iceberg, eroded by waves, Ross Sea, Antarctica. February 2017. Photographed for The Freshwater Project