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Little blue penguin (Eudyptula minor), two standing on rocks at night, silhouetted against Melbourne city lights. St Kilda breakwater, Victoria, Australia. December 2016
Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Rough seas at d'Ar-Men lighthouse during Storm Ruth, Ile de Sein, Armorique Regional Park. Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, September
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Butt of Lewis lighthouse, Port Nis / Port of Ness, Isle of Lewis, Outer Hebrides, Scotland, UK. March, 2015
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Humpback whale (Megaptera novaeangliae) breaching, near Hout Bay, South Africa
Sea stacks, Duncansby Head, John O Groats, Caithness, Scotland, UK, April 2015
Greenland shark (Somniosus microcephalus) with parasitic copepod (Ommatokoita elongata), under ice, Lancaster Sound, Nunavut, northern Baffin Island, Canada, Arctic
Northern gannets (Morus bassanus) diving for fish, split level shot, Shetland, Scotland, UK, July
Sunrise at Rattray Head Lighthouse, north-east Scotland, January 2014. All non-editorial uses must be cleared individually
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Orca whales (Orcinus orca) pod surfacing together, Shetland, Scotland, UK. April
Grey seal (Halichoerus grypus) swimming towards camera, Orkney, Scotland, UK, August
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Weedy seadragon (Phyllopteryx taeniolatus) male carries eggs through a kelp forest (Macrocystis pyrifera) in Tasmania, Australia. Tasmania is the only part of Australia with giant kelp forests
Florida manatee (Trichechus manatus latirostris) mother and calf endangered. Three Sisters Spring in Crystal River, Florida, USA
Gannets (Morus bassanus) diving to feed on discarded fish, Shetland, Scotland, UK, April
Nudibranch (Janolus cristatus) Vela Luka, Korcula Island, Croatia, Adriatic Sea, Mediterranean
RF - Southern stingray (Dasyatis americana) swimming over sand in shallow water at dawn, Cayman Islands, Caribbean Sea
Male Leach's sea star (Leiaster leachi) broadcast spawning, releasing streams of sperm from its arms. Kagoshima Prefecture, Kyushu, Japan
Dense growth of Bootlace seaweed (Chorda filum) in shallow water, Kimmeridge, Dorset, England, UK, English Channel. July. Wildlife Photographer of the Year Competition 2022 (WPY)
Sea otter (Enhydra Lutris) with forelegs raised, Sitka Sound, Alaska, USA, August
RF - Ocellate octopus (Amphioctopus mototi) swimming up from the sea floor. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Great hammerhead shark (Sphyrna mokarran) mouth wide open, feeding in shallow water, surrounded by fish. South Bimini Island, Bahamas
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Indo-Pacific bottlenose dolphin (Tursiops aduncus) with penis extended. Ogasawara / Bonin Islands, Japan
RF- White horse of the Camargue, running from sea. Camargue, Southern France
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Giant Manta Ray (Manta birostris), San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico. Vulnerable species
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Pod of Indian Ocean bottlenose dolphin (Tursiops adunctus) swim over a coral reef. Sha ab El Erg, Hurghada, Egypt. Red Sea
RF - Composite image of tropical nudibranchs on a black background showing variety and abundance of nudibranch species, Indo-Pacific
Deep sea jellyfish (Periphylla periphylla) juvenile, Trondheimsfjord, Norway, Atlantic ocean
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Reflections of clouds in ripples of sea water, Svalbard, Norway, June 2010
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Blue sky reflected in ripples of water, Svalbard, Norway, June 2010
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
Angular roughshark (Oxynotus centrina) a deepsea species living at 80-300m depth, caught by fishermen and released, Costa Brava, Catalunya, Spain, Mediterranean Sea
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Smooth hammerhead shark (Sphyrna zygaena). Cabo San Lucas, Baja California Sur, Mexico