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Brown bear (Ursus arctos) catching salmon in river, Kamchatka, Far east Russia, August
Female Atlantic salmon (Salmo salar) River Orkla, Norway, October 2008
Atlantic salmon (Salmo salar) migrating upstream to spawn, Umelven, Sweden, July 2009
Captured Atlantic salmon (Salmo salar) awaiting tagging by Tweed Foundation staff monitoring fish movement, density and biology, River Tweed, Berwickshire, Scotland, UK, September
Atlantic salmon (Salmo salar) leaping up the cauld at Philphaugh Salmon Viewing Centre near Selkirk, where members of the public can view the spectacle, Selkirkshire, Scotland, UK, October 2011
Atlantic Salmon (Salmo salar) in pool on upstream spawning migration, silhouetted against sky. Quebec, Canada, August
Sockeye salmon (Oncorhynchus nerka) split level view of annual spawning run, Adams River, British Columbia, Canada, October
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
Atlantic salmon (Salmo salar) leaping upstream in river, Scotland, UK
Brown bear / Coastal Grizzly Bear, (Ursus arctos) feeding on Pink Salmon, during the autumn salmon run. Hallo Bay, Katmai National Park, Alaska, USA
Salmon (Salmonidae) fry with egg case still attached, Lake Kuril, Kamchatka, Far East Russia, January
Looking up at gills of Atlantic salmon (Salmo salar) River Orkla, Norway, September 2008
Atlantic salmon (Salmo salar) River Orkla, Norway, September 2008
Atlantic salmon (Salmo salar) male, River Orkla, Norway, September 2008
Juvenile Atlantic salmon (Salmo salar) fry, River Orkla, Norway, September 2008
Atlantic salmon (Salmo salar) female, River Orkla, Norway, September 2008
Atlantic salmon (Salmo salar) just below the surface, Orkla River, Norway, September 2008
Grizzly bears (Ursus arctos horribilis) looking for salmon, Brooks river, Katmai national park, Alaska, North America
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July