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Chilean flamingos (Phoenicopterus chilensis) in flight over mountain peaks with glacier in the distance, Torres Del Paine National Park, Chile
Whale Shark (Rhincodon typus) passing by baitball, Santa Maria Island, Azores, Portugal, September 2008 Exclusive Japanese calendar rights for 2014
Large shoal of Horse mackerel (Megalaspis cordyla) schooling. Rinca, Komodo National Park, Indonesia
Aggregation of thousands of Spider crabs (Leptomithrax gaimardii) for moulting, South Australia Basin, Australia. Pacific Ocean
School of large Atlantic bluefin tuna (Thunnus thynnus) captive in growing pen. Depsite looking much smaller in the photo, these fish were 1.5m (5ft) long. Malta. Mediterranean Sea
Lesser flamingo flock (Phoeniconaias) flying over lake with green algae, aerial view, Rift valley, Kenya
Thousand of Gilt-head bream (Sparus aurata) inside a sea cage used for aquaculture, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean
Cheetah (Acinonyx jubatus) cubs 4 months, Masai-Mara Game Reserve, Kenya. Vulnerable species
A large herd of Tien Shan Argali (Ovis ammon karelini) making tracks through snow. Naryn National Park, Kyrgyzstan, Central Asia, November
Waterfowl at dawn with mist, including Great blue heron (Ardea herodias herodias), American avocet (Recurvirostra americana), Northern shoveler (Anas clypeata), Xochimilco wetlands, Mexico City
Dalmatian pelicans (Pelecanus crispus) group on water in mist, Lake Kerkini, Greece, February
Coral reef scenery with Gorgonian and shoal of Pygmy / Glassy sweepers (Parapriacanthus guentheri) Egypt, Red Sea
Coalition of male Lions (Panthera leo) in grassland before storm, Masai-Mara game reserve, Kenya
Whale shark (Rhincodon typus) with shoal of anchovies staying near to avoid predation. Los Roques, Venezuela. Caribbean
Four Cedar waxwings (Bombycilla cedrorum) feeding on Crabapple (Malus sp) fruit, Ithaca, New York, USA, February
Turkmenian Wild Ass / Kulan (Equus hemionus kulan) Badkhyz, Turkmenistan. Endangered
A split level photo of group of Sockeye salmon (Oncorhynchus nerka) fighting their way upstream as they migrate back to the river of their birth to spawn, trees showing autumnal colours
RF- Elongate surgeonfish (Acanthurus mata) school. Indonesia
Large school of Bohar snappers (Lutjanus bohar) gather with smaller shoal of Batfish (Platax orbicularis) in open water off Ras Mohammed at the tip of Sinai, Egypt
Herd of Tibetan Wild Asses / Kiang (Equus kiang) ChangThang, Tso Kar lake, at altitude of 4600m, Ladakh, India, October 2012
Wildebeest (Connochaetes taurinus) crossing the Mara river, Masai-Mara Game Reserve, Kenya
Rothschilds giraffe (Giraffa camelopardalis rothschildi), Murchison Falls National Park, Uganda, East Africa. Murchison Falls National Park hosts the only remaining naturally occurring population of
A flock of Greylag Goose (Anser anser) flying in formation above misty fields. The Netherlands, March
Little Auks (Alle alle) flying low above surface in front of iceberg. Spitsbergen, Svalbard, Norway, July 2010. Highly commended in the Birds category in the GDT competition 2011
Bigeye Scad (Selarcrumenophthalmus). Misool, Raja Ampat, West Papua, Indonesia
Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December
Smack of Moon jellyfish (Aurelia aurita), Prince William Sound, Alaska, USA, Pacific Ocean
European Spider crab aggregation (Maja squinado) St. Ives, Cornwall. AugustEuropean Spider crab aggregation (Maja squinado) St.Ives, Cornwall. August
RF- Caribou herd (Rangifer tarandus) grazing on tundra. Kobuk Valley National Park, Alaska, USA
Oriental sweetlips (Plectorhinchus vittatus), Maldives, Indian Ocean
Fulmars (Fulmarus glacialis) on water, feeding at a fish processing plant outfall. Grundafjordur, Iceland, March
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Lyretail anthias / Goldies (Pseudanthias squamipinnis) over coral reef with soft corals (Dendronephthya sp) Egypt, Red Sea
Sockeye / Red Salmon (Oncorhynchus nerka) on spawning migration. Adams River, British Columbia, Canada, October
School of Bohar snappers (Lutjanus bohar) Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Shoal of Longfin bannerfish (Heniochus acuminatus) Maldives, Indian Ocean
Shoaling glassfish (Parapriacanthus sp. ) shoal, Maldives, Indian OceanShoaling glassfish (Parapriacanthus sp.) shoal, Maldives, Indian Ocean
Snow Geese (Chen caerulescens) in flight, silhouetted against colourful dusk sky. Bosque del Apache, New Mexico, USA, November
Monarch butterfly (Danaus plexippus) hibernating, Mariposa Monarca Special Biosphere Reserve, central Mexico, January
American Flamingo (Phoenicopterus ruber), Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, August
A school of lesser devil rays (Mobula hypostoma) flying through sunbeams as they feed. Isla Mujeres, Quintana Roo, Yucatan Peninsula, Mexico. Caribbean Sea
White terns (Gygis alba) flock in flight overhead, Christmas Island / Kiritimati, Pacific Ocean, July
Flock of Greater flamingo (Phoenicopterus ruber) in flight, Camargue, Southern France, November 2006
Mute swans (Cygnus olor) newly hatched cygnets on pond, Yetholm Loch Scottish Wildlife Trust Reserve, Roxburghshire, Scotland, May
Row of Starlings (Sturnus vulgaris) perched on wire. Marais Breton, Vendee, French Atlantic Coast, November
Wild / feral Dulmen ponies (Equus caballus) close up of herd of mares and foals running on the Duke of Croys estate, Meerfelder Bruch, North Rhine-Westphalia, Germany, May 2011
Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October. Non-ex
Black-legged Kittiwake (Rissa tridactyla) on ice. Floe edge, Arctic Bay, Nunavut, Canada, April