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1000 year old grove of Banyan fig trees (Ficus benghalensis) Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan province, China, May 2017
English oak tree (Quercus robur) trunk in winter, Hampstead Heath, London, February
Lemon shark juvenile (Negaprion brevirostris) trying to feed on the leaves of a red mangrove (Rhizophora mangle). As pups, the sharks have to learn to hunt for themselves and sometimes make mistakes
Spotted eagle rays (Aetobatus narinari) shoal among mangrove roots, Elizabeth Bay, Isabela Island, Galapagos
Yellow Anaconda (Eunectes notaeus) in vegetation at the edge of the Paraguay River, Taiama Reserve, western Pantanal, Brazil, South America
Sergeant major (Abudefduf sp) schooling in mangroves of Isabela Island, Galapagos
Strangler fig (Ficus sp) growing over Hoop Pine tree(Araucaria cunninghamii) with branches knotting round each other, Morans Falls Track, Green Mountains
Jaguar (Panthera onca palustris), male hunting amongst tree roots at edge of the Cuiaba River. Three Brothers, Porto Jofre, northern Pantanal, Mato Grosso, Brazil. August 2017
Tea mangroves (Pelliciera rhizophorae) Pochote Estuary, Costa Rica. Vulnerable species
Beech (Fagus sylvatica) mature tree in winter, Paxton House Estate, Berwickshire, Scotland, December
Purple gallinule (Porphyrula martinica) in mangrove swamp. Everglades National Park, Florida, USA. March
Great Basin Bristlecone Pine (Pinus longaeva) fallen ancient tree, White Mountains, California, USA, March
Tropical forest, Andasibe-Mantadia National Park, Madagascar, March 2005
Sika deer (Cervus nippon) herd feeding beside pool in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Water lettuce (Pistia stratiotes) with submerged roots in aquarium
Lemon shark (Negaprion brevirostris) pup and school of fish swimming through Red mangrove (Rhizophora mangle) nursery, Eleuthera Island, Bahamas, North Atlantic
Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp)Orbiculate Cardinalfish (Sphaeramia orbicularis) school sheltering amongst roots of mangroves (Rhizophora sp.) Gam Island, Raja Ampat Islands, West Papua, Indonesia
Ocelot (Leopardus pardalis) hiding amongst tree roots, Costa Rica, Central America, 2016. Filmed for the BBC series Big Cats
Laurisilava forest. Natural Monument of Caldeira Velha, Ribeira Grande, Sao Miguel Island, Azores, Portugal
Needle bug nymph (Ranatra linearis) hidden in roots and waiting for prey, Europe, July, controlled conditions
Rhinoceros beetle (Chalcosoma moellenkampi) climbing up tree trunk, Danum Valley Reserve, Sabah, Borneo, Malaysia
Hanuman / Northern Plains Grey Langur (Semnopithecus entellus) youngster sitting on the trunk of a Banyan tree, Ranthambore National Park, Rajasthan, India, June
Looking through the aerial roots of a Strangler fig (Ficus zarazalensis) the original host tree that the Fig was climbing has been killed and has rotted away, leaving a hollow space in the middle