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Killer Whales / Orcas (Orcinus orca)Killer whales / Orcas (Orcinus orca) large pod including calf traveling together while foraging on large schools of Herring (Clupea harengus) in the cold waters of northern Norway, January
RF - Hawksbill sea turtle (Eretmochelys imbricata) swimming over a coral reef. Whilst a Predatory bandcheek wrasse (Oxycheilinus digrammus)
Piraputanga fish (Brycon hilarii) in underwater landscape, Aquario Natural, Rio Baia Bonito, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso do Sul, Brazil
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
Northern pike (Esox lucius) amongst Shining pondweed (Potamogeton lucens). Lake Neuchatel, Switzerland, September. Photographed for The Freshwater Project
Tarpon (Megalops atlanticus) portrait, Eleuthera, Bahamas. December
Dugong (Dugong dugon) male feeding on a seagrass meadow (Halophila stipulacea), accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
School of Blackfin barracuda (Sphyraena qenie) forming circle in open water off the wall at Shark Reef, Ras Mohammed Marine Park, Sinai, Egypt. Red Sea
Weedy seadragon (Phyllopteryx taeniolatus) male carries eggs through a kelp forest (Macrocystis pyrifera) in Tasmania, Australia. Tasmania is the only part of Australia with giant kelp forests
Tompot blenny (Parablennius gattorugine) portrait, Swanage Pier, Swanage, Dorset, UK, English Channel, August
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Kaleidoscopic image of Clarks anemonefish (Amphiprion clarkii), Ambon, Indonesia
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
A portrait of a Mediterranean parrotfish (Sparisoma cretense) above an algae covered rock. El Hierro, Canary Islands, Spain. East Atlantic Ocean
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Illuminated netdevil (Linophyne arboritera), male anglerfish attached to much larger female, captive from deep sea
Piraputanga (Brycon hilarii) shoal, Rio Olhio d agua, tributary of Rio da Prata, Bonito area, Serra da Bodoquena (Bodoquena Mountain Range), Mato Grosso del Sul, Brazil
Pletail unicornfish (Naso brevirostris) juvenile, its horn will continue to grow as it gets older. Fiji Pacific Ocean
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Backlit photo of a Robust ghost pipefish (Solenostomus cyanopterus) reveals a small fish that it has just eaten in its stomach. Lembeh Strait, North Sulawesi, Indonesia. Molluca Sea
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Turbot (Scophthalmus maximus) detail of the eyes and mouth, United Kingdom, September
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
West African mudskipper (Periophthalmus barbarus) Joao Vieira Poillo, Bijagos Archipellago. Guinea Bissau
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Short-snouted seahorse (Hippocampus hippocampus) male, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean Sea
Giant hogfish (Bodianus macrognathos) male yawning at a cleaning station run by juvenile Spanish hogfish (Bodianus rufus). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands
Unicorn leatherjacket filefish (Aluterus monoceros) at night, The Bahamas. February
Tiger moray / Fangtooth Moray (Enchelycore anatina) and Black moray eel (Muraena augusti), South Tenerife, Canary Islands, Atlantic Ocean
Couple of Blue shark (Prionace glauca) and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Blue shark (Prionace glauca) with fishing hook caught in mouth and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands