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RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
RF - Marabou (Leptoptilos crumenifer) with Whitebacked vultures (Gyps africanus) Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Leaves of a Ginkgo tree or Maidenhair tree (Ginkgo biloba) Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Red-billed blue magpie (Urocissa erythrorhyncha) perched on branch, Tangjiahe National Nature Reserve, Qingchuan County, Sichuan province, China
Amazon kingfisher (Chloroceryle amazona) with fish, Cuiaba, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) male standing in cold and rain with strong wind blowing, smelling the air, Masai Mara National Reserve, Kenya
Female Bornean orangutan (Pongo pygmaeus) sitting in a tree, Tanjung Puting National Park, Borneo-Kalimantan, Indonesia
Black rhinoceros (Diceros bicornis) covered in mud, Mud on skin Etosha National Park, Namibia
Mountain gorilla (Gorilla beringei beringei) female resting with her baby, members of the Humba group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Mountain gorillas (Gorilla gorilla beringei) grooming, members of the Munyaga group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Moutain gorilla (Gorilla beringei beringei) close up of hand, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Portrait of male silverback Mountain gorilla (Gorilla beringei beringei) Virunga National Park, Democratic Republic of Congo
Eastern lowland gorilla (Gorilla beringei graueri) silverback named Chimanuka, Kahuzi-Biega National Park, South Kivu Province, Democratic Republic of Congo (DRC)
Polar bear (Ursus maritimus) female coming out the den. Wapusk National Park, Churchill, Manitoba, Canada, March
Polar bear (Ursus maritimus) sow standing along snow covered Bernard Spit, off the 1002 Area, Arctic National Wildlife Refuge, North Slope, Alaska, USA, October. Vulnerable species
Frozen icicles and frozen waterfall, Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, April
Northern lights over Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Waterfall in Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia, June 2016
River and cliffs in Putoransky State Nature Reserve, Putorana Plateau, Siberia, Russia
Arctic fox (Vulpes lagopus) in winter fur, licking nose, Wrangel Island, Far Eastern Russia, October
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
Polar bear (Ursus martimus) footprints in snow, Wrangel Island, Far Eastern Russia, March
Polar bear (Ursus maritimus) female with young, age one year and a half, resting on the ice, Wrangel island, Far East Russia
Reindeer (Rangifer tarandus) bull in snow, Cairngorms Reindeer Herd, reintroduced to Scottish Highlands, Cairngorm National Park, Speyside, Scotland, December
Wild Rapa Nui horse / stallion, Rapa Nui National Park UNESCO World Heritage Site, Easter Island / Rapa Nui, Chile
RF - Wild pinto Mustang stallion running in snow, Black Hills Wild Horse Sanctuary, South Dakota, USA. January
Painted stork (Mycteria leucocephala), resting on nest with young at sunrise. Keoladeo / Bharatpur National Park UNESCO World Heritage Site, India
Jungle cat (Felis chaus), female in search of prey. Velavadar National Park, Gujarat, India
Sloth bear (Melursus ursinus) close up portrait, Daroiji Bear Sanctuary, Karnataka, India. Remote camera image
Sloth bear (Melursus ursinus) mother with cub on her back, Daroiji Bear Sanctuary, Karnataka, India
Sloth bear (Melursus ursinus) cubs riding on mothers back, Daroiji Bear Sanctuary, Karnataka, India
Indian wild ass (Equus hemionus khur), lone stallion standing, Little Rann of Kutch, Gujarat, India
Asiatic wild buffalo (Bubalus arnee) portrait of female. Kaziranga National Park, India
Common hoopoe (Upupa epops) sitting on a stone, Tangjiahe National Nature Reserve, Sichuan Province, China
Golden snub-nosed monkey (Rhinopithecus roxellana) hanging from branches, Foping Nature Reserve, Shaanxi, China. Endangered species
1000 year old grove of Banyan fig trees (Ficus benghalensis) Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan province, China, May 2017
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Massive old growth oak trees of the cloud forest, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica. Small repro only
Mountain landscape of Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fossil leaves of seed ferns, a kind of extinct plant, Joggins Fossil Cliffs UNESCO World Heritage Site, Bay of Fundy, Nova Scotia, Canada. May 2017
Semipalmated sandpiper (Calidris pusilla) flock in flight over water, Johnsons Mills Reserve, Bay of Fundy, New Brunswick, Canada, July