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Mountain hare (Lepus timidus) in summer pelage amongst heather, Cairngorms National Park, Scotland, UK, August
Curled octopus (Eledone cirrhosa) swimming over sea floor, South Arran Marine Protected Area, Isle of Arran, Scotland, UK, August
Peacock butterfly (Inachis io) feeding on Field Scabious (Knautia arvensis) in a disused limestone quarry. Peak District National Park, Derbyshire, UK. August
Ash Tree (Fraxinus excelsior) growing in a field. Peak DIstrict National Park, Derbyshire, UK. August
Tufted duck (Aythya fuligula) male drake, Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Great white egret (Egretta Alba) preening its feathers, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, July
Great crested grebe (Podiceps cristatus) adult with young on its back, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Mute swan (cygnus olor) family, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, June
Alpine ibex (Capra ibex) adult males fighting in alpine landscape, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Alpine ibex (Capra ibex) struggling in deep snow on a steep slope, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, January
Tayra (Eira barbara) climbing in a tree in rainforest habitat with fern covered trees, Bellavista private reserve, Mindo cloud forest area, Ecuador, July
Mountain gorilla (Gorilla beringei) juvenile aged 2 years, hanging from branch, member of the Nyakagezi group, Mgahinga National Park, Uganda. January
Phuket horned tree lizard (Acanthosaura phuketensis) near Bang Pae Waterfall, Khao Phra Thaew NP, Phuket Island, Thailand
Autumnal beech trees ((Fagus sylvatica) at Bolderwood, The New Forest, Hampshire, UK. November
Clapper Bridge at Postbridge, early morning light and mist, Dartmoor National Park, Devon, UK. October 2018
Black-necked stilts (Himantopus mexicanus) pair performing post-copulatory display (as copulation ends, the male slides off the female leaving his wings draped over her)
Black-necked Stilt (Himantopus mexicanus), foraging in water, with reflection, Bear River Migratory Bird Refuge, Utah, USA, May
European otter (Lutra lutra) two at waters edge, one shaking off water, Kiskunsagi National Park, Pusztaszer, Hungary, October
European otters (Lutra lutra) feeding at water?s edge with splashing water, Kiskunsagi National Park, Hungary, January
Lone Scots pine tree (Pinus sylvestris) and star trails with the north star, Abernethy forest, Cairngorms National Park, Scotland, UK
Asian black bear (Ursus thibetanus) feeding on leaves, Tangjiahe Nature Reserve, Sichuan Province, China, April
Tibetan wolf (Canis lupus chanco) in mountain landscape, Serxu County, Garze Prefecture, Sichuan Province, China
Snow leopard (Uncia uncia) with Bharal (Pseudois nayaur) prey, Serxu County, Garze Prefecture, Sichuan Province, China
Tibetan macaque (Macaca thibetana) carrying young baby, Tangjiahe Nature Reserve, Sichuan Province, China
Sumatran Orangutan (Pongo abelii) female with very young baby, Gunung Leuser National Park, Sumatra, Indonesia
Himalayan honey bees (Apis dorsata laboriosa) on comb, Mount Namjagbarwa, Yarlung Zangbo Grand Canyon National Park, Tibet, China. October
Pine marten (Martes martes) foraging in Pine (Pinus sp) woodland at night. Glenfeshie, Cairngorms National Park, Scotland, UK
Osprey (Pandion haliaetus) fishing. Rothiemurchus, Cairngorms National Park, Scotland, UK
Ancient Scots pine (Pinus sylvestris) amongst flowering Common heather / Ling (Calluna vulgaris). Rothiemurchus Forest, Cairngorms National Park, Scotland, UK
Bigeye trevally (Caranx sexfasciatus), a large shoal. Cocos Island National Park, Costa Rica
Eurasian badger (Meles meles), two foraging in Pine (Pinus sp) woodland. Glenfeshie, Cairngorms National Park, Scotland, UK
Rocks and coastal rainforest. Cocos Island National Park, Costa Rica. 2018
Squirrelfish (Holocentrus sp) shoal amongst rocks. Near Cocos Island National Park, Costa Rica
Blue-lined snapper (Lutjanus kasmira) shoal. Cocos Island National Park, Costa Rica
Fish seeking shelter around floating tree in open ocean. Off coast of Cocos Island National Park, Costa Rica
Central American spider monkey (Ateles geoffroyi) juveniles hanging by tails and playing, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Muskox (Ovibos moschatus), male portrait. Dovrefjell National Park, Norway
Cuban crocodile (Crocodylus rhombifer) in a cenote in Cienaga de Zapata National Park. Cuba. Critically endangered species
Male American Bison (Bison bison) walking through snow. Firehole River Valley, Yellowstone National Park, Wyoming, USA. January
Trumpeter swans (Cygnus buccinator) on the edge of the Upper Yellowstone River. Hayden Valley, Yellowstone National Park, Wyoming, USA. January
Australian sea lion (Neophoca cinerea), two females greeting each other on beach. Seal Bay Conservation Park, Kangaroo Island, South Australia
One-horned Asian rhinoceros (Rhinoceros unicornis), Chitwan National Park, Inner Terai lowlands, Nepal
Arctic fox (Vulpes lagopus), juvenile licking lips, winter pelage. Dovrefjell National Park, Norway. September
Indian rhinoceros (Rhinoceros unicornis) Kaziranga National Park, Assam, India
Melanistic leopard / Black panther (Panthera pardus) Nagarahole National Park, Nilgiri Biosphere Reserve, Karnataka, India
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
American crocodile (Crocodylus acutus) rear view of animal resting in shallow water with head at surface, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico
Red fox (Vulpes vulpes) resting with eyes open, Vanoise National Park, Rhone Alpes, France. October