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Single Male waterbuck (Kobus ellipsiprymnus) in distance silhouetted against big sky and clouds, Masai-Mara Game Reserve, Kenya. October
Female Persian leopard (Panthera pardus saxicolor), captive, occurs in the Caucasus, Turkmenistan and Afghanistan
Leaf tailed gecko (Uroplatus phantasticus) concealed in amongst dry leaves, Madagascar
Profile portrait of Spotted hyaena (Crocuta crocuta) Moremi WR, Okavango Delta, Botswana
Herons at night on Lake Csaj, Kiskunsag National Park, Hungary. Winner of the Birds category, Wildlife Photographer of the Year (WPOY) Competition 2014
Hawksbill turtle (Eretmochelys imbricata) Port El Ghalib, Egypt, Red Sea
Impala (Aepyceros melampus) male, Masai-Mara Game Reserve, Kenya
Mountain Clouded Yellow (Colias phicomone) profile portrait, Aosta Valley, Monte Rosa Massif, Pennine Alps, Italy
Golden Eagle (Aquila chrysaetos) portrait in snow, Utajarvi, Finland February
Red Squirrel (Sciurus vulgaris) adult reaching up. Cairngorms National Park, Scotland, UK, March
Dall sheep (Ovis dalli) solitary animal at sunset with a storm moving across the sky, Denali National Park, interior Alaska, USA
Salmon landing on head of Grizzly bear (Ursus arctos horribilis) as it is leaping up rapids, Katmai National Park, Alaska, USA
Spectacled caiman (Caiman yacare) walking on a sandy shore at the edge of a river at night, Pantanal, Brazil
Little egret (Egretta garzetta) attempting to catch fish, Camargue, France, July
Whiskered auklet (Aethia pygmaea), Iony Island / Jonas Island, Sea of Okhotsk, Far East Russia, July
Black faced Impala (Aepyceros melamis petersi) female jumping, Etosha National Park, Namibia
Mistle Thrush (Turdus viscivorus) in snow. Hertfordshire, England, UK, February
Southern plains grey langur ( Semnopithecus entellus / Presbytis entellus) close up face profile, Sawai Modhopur, Rajasthan, India
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Cotswolds Lion rare breed sheep (Ovis aries) and the village of Naunton at sunset, Gloucestershire, UK, June 2001
Red Grouse (Lagopus lagopus scoticus) in Heather. Peak District, UK, August
Two Least Chipmunks (Tamias minimus) sitting on tree stump feeding, Yellowstone National Park, Wyoming, USA, October, crop of 1373882
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Caribbean reef shark (Carcharhinus perezi) Jardines de la Reina National Park, Cuba
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Red squirrel (Sciurus vulgaris) standing on its hind legs, Cairngorms National Park, Scotland, UK, March. Non-ex
Black-Winged Stilt (Himantopus himantopus) wading. Etosha National Park, Namibia, January
Jaguar (Panthera onca palustris) male wading through water at the shore of the Piquiri River. The Pantanal wetlands of Mato Grosso, Mato Grosso State, Brazil, October
A juvenile Weedy Seadragon (Phyllopteryx taeniolatus) in its habitat. Flinders Jetty, Melbourne, Victoria, Australia, March
Long Water Measurer (Hydrometra stagnorum) drinking by piercing surface tension with mouthparts. Captive, UK, July
Blue Tit (Parus caeruleus) perched on mossy branch. Breton Marsh, French Atlantic Coast, November
Polar Bear (Ursus maritimus) walking on icepack with arctic coast in the background. Floe edge, Arctic Bay, Baffin Island, Nunavut, Canada, April
Chestnut Paso peruvian stallion running, California, USA
Yellow Labrador Retriever standing in glasswort against a blue sky. Monterey Peninsula, California, USA, February
Portrait of a wild Horse (Equus caballus). The foothills of southwest Reno, Nevada, USA, May
Portrait of an Arctic snow hare (Lepus arcticus groenlandicus) North East Greenland. March
African lion (Panthera leo) male and female at sunrise, Masai Mara National Reserve, Kenya, January
Red grouse {Lagopus lagopus scoticus} on heather moorland, Derwent Edge, Peak District National Park, UK, September
Red Squirrel (Sciurus vulgaris) leaping through Ash Tree in deciduous woodland, Berwickshire, Scotland, UK, February
Little egret {Egretta garzetta} at waters edge hunting sequence 1 / 3, Donana NP, SevillaLittle egret {Egretta garzetta} at waters edge hunting sequence 1/3, Donana NP, Sevilla, Spain
African fish eagle about to catch fish {Haliaeetus vocifer} Chobe NP, Botswana
Corncrake (Crex crex) calling from rock, North Uist, Outer Hebrides, Scotland, UK
Dall sheep {Ovis Dalli}female profile walking on hillside, Denali National Park, Alaska, USA
Female European Brown bear with cubs following {Ursus arctos} Lapland Finland
Solitary Leopard {Panthera pardus} walking along sand dune Namibia, Southern Africa
Marine iguana {Amblyrhynchus cristatus} Fernandina Is, Galapagos Islands, Ecuador
RF - Chinstrap Penguins (Pygoscelis antarcticus) once calling, South Shetland Islands, Antarctica Continent