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Blue whale (Balaenoptera musculus) diving beneath ocean surface. Indian Ocean, Sri Lanka
Dartford warbler (Sylvia undata) singing on Gorse, Dorset, UK, April
Barn owl (Tyto alba) in flight, hunting, Hampshire, England, UK. Captive
Mountain Gorilla (Gorilla beringei beringei) Sabyinyo Group, Silverback in meadow, Volcanoes National Park, Rwanda, East Africa
Giant pangolin (Manis gigantea) walking along ground, Tanzania
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Polar bear (Ursus maritimus) young bear rolling around in the snow, on newly formed pack ice during autumn freeze up, Beaufort Sea, off Arctic coast, Alaska
Bottlenose dolphin (Tursiops truncatus) two breaching in evening light, Moray Firth, Inverness-shire, Scotland, UK, August, sequence 2/2
Head portrait of black Merens stallion with long mane running in pasture. Northern France, Europe. February
Portrait of a Mandarin duck (Aix sponsa) male animal and female. UK. Introduced species
Glanville fritillary butterfly (Melitaea cinxia) roosting on larval foodplant Ribwort plantain (Plantago lanceolata), UK - Captive - Focus Stacked Image
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Adelie penguin (Pygoscelis Adeliae) in falling snow on the western Antarctic Peninsula, Southern Ocean
Giant anteater (Myrmecophaga tridactyla) Serra de Canastra National Park, Brazil
Shoebill stork (Balaeniceps rex) in the swamps of Mabamba, Lake Victoria, Uganda
Killer whale (Orcinus orca) pod hunting together in herring baitball (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
A portrait of a male Ballan wrasse (Labrus bergylta), showing his bright mating colours. Plymouth, Devon, England, Great Britain. Wreck of the Glen Strathallan, Plymouth Sound, English Channel
Head portrait of grey Arabian stallion running in desert dunes near Dubai, United Arab Emirates
House Martin (Delichon urbica) Spain, April
Arctic fox (Vulpes lagopus) portrait of juvenile, winter pelage. Dovrefjell National Park, Norway. September
Arctic fox (Vulpes lagopus) juvenile sniffing ground, winter pelage. Dovrefjell National Park, Norway. September
Male Black rhinoceros (Diceros bicornis) running, Nakuru National Park, Kenya
Mallard (Anas platyrhynchos) male flying low over water. Cambridgeshire Fens, England, March
Red deer (Cervus elaphus) stag silhouetted at sunset, Scotland, UK, February
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Elephant hawk moth (Deilephila elpenor) portrait, Brackagh Moss National Nature Reserve, Portadown, County Armagh, Northern Ireland. April
Grass Snake (Natrix natrix) portrait with tongue extended, Staffordshire, England, UK, April
Fluffy black kitten, 12 weeks old, stretching
Silver Studded Blue (Plebejus argus), Gran Paradiso National Park, Aosta Valley, Pennine Alps, Italy. July
Atlantic Puffins (Fratercula arctica) pair bill rubbing, part of ritual courtship, Skomer Island, Wales, UK, May
Eurasian blue tit (Cyanistes caeruleus) perched in tree amongst catkins. Denmark, February
Jay (Garrulus glandarius) perched in Oak tree in autumn, Hertfordshire, England
Black crowned-crane (Balearica pavonina) foraging in wetland area, Masai Mara, Kenya
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Oystercatcher (Haematopus ostralegus) by the coast, Island of Runde, Norway July
Mosquito (Aedes punctor) female resting after sucking blood from human arm. Sequence 4 / 4Mosquito (Aedes punctor) female resting after sucking blood from human arm. Sequence 4/4
Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Black-throated Diver (Gavia arctica) Rokua, Finland, May
RF - Bison (Bison bison) breathing in the cold air, Yellowstone National Park, USA, February
Giant monolithic stone Maoi statues at Rano Raraku, Easter Island, Rapa Nui, Chile, South America 2008
RF- African elephant (Loxodonta africana) at sunset. Chobe National Park, Botswana. Endangered species
Portrait of a Black-capped chickadee, (Poecile atricapillus) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Wild Eurasian lynx (Lynx lynx) at night with city lights and sky glow behind, Switzerland. December. Taken with remote camera trap
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May