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RF- Portrait of three kittens
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Arctic fox (Vulpes lagopus) in winter fur, licking nose, Wrangel Island, Far Eastern Russia, October
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
Siberian tiger (Panthera tigris altaica) in snow, captive
Red squirrel (Sciurus vulgaris) stood on log in snow, Cairngorms National Park, Scotland, UK. December
RF - Head portrait of grey Andalusian mare with long mane in snow, Berthoud, Colorado, USA. January
Portrait of Skewbald Icelandic horse in the snow, Snaefellsnes Peninsula, Iceland, March
Exmoor ponies (Equus caballus) in snow, Aurochs breeding site run by The Taurus Foundation, Keent Nature Reserve, The Netherlands
Young girl wearing traditional headdress, Mursi tribe, Mago National Park. Omo Valley, Ethiopia
Jungle cat (Felis chaus), female in search of prey. Velavadar National Park, Gujarat, India
Sloth bear (Melursus ursinus) close up portrait, Daroiji Bear Sanctuary, Karnataka, India. Remote camera image
White-lipped pit viper (Trimeresurus albolabris) Lantau Island, Hong Kong, China
Tokay gecko (Gekko gecko) Shek Pik, Lantau Island, Hong Kong, China
Stellar rockskipper (Entomacrodus stellifer) hiding in a barnacle. Sai Kung East Country Park, Hong Kong, China
Glovers pika (Ochotona gloveri), Baixha Valley, Tibetan Plateau, Qinghai, China
Atlantic wolfish (Anarhichas lupus) looking out hole, Bay of Fundy, New Brunswick, Canada, July
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Reed warbler (Acrocephalus scirpaceus) with insect larvae in beak, Greylake RSPB Reserve, Somerset Levels, England, UK, June
RF - Snow leopard (Panthera uncia) portrait, captive
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Vicuna (Vicugna vicugna) Uyuni, Altiplano, Bolivia
Komodo dragons (Varanus komodoensis) on shore with saliva dripping from mouth, Rinca Island, Komodo National Park, Indonesia
Snow leopard (Panthera uncia) cub age three months, captive
Cliff parakeets (Myiopsitta luchsi) perched on cactus, Red-fronted Macaw Community Nature Reserve, Omerque, Bolivia
Southern stingray (Dasyatis americana) large female foraging over seagrass in shallow water. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Coral Rabbitfish (Siganus corallinus) portrait, Tubbataha Reef Natural Park, UNESCO World Heritage Site, Sulu Sea, Cagayancillo, Palawan, Philippines
Pallass cat (Otocolobus manul) two sitting side by side, Tibetan Plateau, Qinghai, China
Highland Cow (Bos taurus) close-up, Isle of Mull, Inner Hebrides, Scotland, April
Beautiful pit viper (Trimeresurus venustus) captive occurs in Thailand
Green and black poison dart frog (Dendrobates auratus), La Selva Field Station, Costa Rica
Peacock mantis (Pseudempusa pinnapavonis) in defensive posture; captive occurs in Burma and Thailand
RF - Thistle Mantis (Blepharopsis mendica) among petals, captive, occurs in North Africa and the Canary Islands
East Caucasian tur (Capra cylindricornis) portrait, Caucasus, Russia. May
Amur leopard (Panthera pardus orientalis) Land of the Leopard National Park, Primorsky Krai, Far East Russia. March
Close up of a juvenile Pallas cat (Otocolobus manul) resting in its den, Mongolia, June
Green huntsman spider (Olios sp), Intervales State Park, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Portrait of a Golden snub-nosed monkey 1+Rhinopithecus roxellana+2 full frame of the face, Foping Nature Reserve, Shaanxi, China. Endangered species
Shoebill (Balaeniceps rex) portrait. Swamps of Mabamba, Lake Victoria, Uganda
RF - Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Mountain gorilla (Gorilla beringei beringei) silverback male, portrait, member of the Nyakagezi group, Mgahinga National Park, Uganda. Critically endangered
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Blue shark (Prionace glauca) off Halifax, Nova Scotia, Canada. July
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Reindeer (Rangifer tarandus) female from reintroduced herd on Cairngorm Mountains, Cairngorm National Park, Speyside, Scotland, December
European otter (Lutra lutra) with snow ball, Bavarian Forest National Park, Germany, January. Captive
