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European honey bee (Apis mellifera) feeding on ice plant flowers (Sedum spectabile), Monmouthshire, Wales, UK. September
Golden eagle (Aquila chrysaetos) perched in snow covered tree. Kalvtrask, Vasterbotten, Lapland, Sweden. January
Freshwater fishes including Cichlid between water plants and leaves. Cenote Nicte-Ha, near Tulum, Quintana Roo, Yucatan Peninsula, Mexico
Fiordland crested penguins (Eudyptes pachyrhynchus) in dense coastal forest, Lake Moeraki, South Island, New Zealand, November, threatened species
Bearded tit (Panurus biarmicus) male portrait feeding, Helsinki, Finland, January
Puffin (Fratercula arctica) on a cliff edge with flowering sea thrift (Armeria maritima) Great Saltee, Saltee Islands, County Wexford, Ireland, June
Rainbow at Iguazu Falls, Brazil / Argentina border. Photographed from Argentina. August 2017
Line of Leaf-cutter ants (Atta sp) carrying leaves, digital composite
Candover Brook, tributary of the River Itchen, chalk stream Hampshire, England, UK, June. Photographed for The Freshwater Project
RF - Red Deer stag (Cervus elaphus) in snow-covered pine forest. Scotland, UK. December
RF - Red Squirrel (Sciurus vulgaris) on log in snow. Scotland, UK. December
Fieldfare (Turdus pilaris) in Rowan tree with berries in winter, Jyvaskyla, Central Finland, January
Star trails above Scots pine (Pinus sylvestris) woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, March 2017
Scots pine (Pinus sylvestris) trees reflected in Loch Maree at dawn with Slioch in background, Wester Ross, Scotland, UK, November 2014
Pine marten (Martes martes) climbing Scots pine tree, Scotland, UK. May
Water lilies (Nymphaeaceae) seen from below, Cenote Car Wash / Aktun Ha, Mexico. Yucatan peninsula
Tadpoles of the Common toad (Bufo bufo) swimming seen from below, Belgium, June
Darwins Orchid (Angraecum sesquipedale) species which is pollinated by a long-tongued moth, from Ambila, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, August 2009
Bamburgh Castle and sand dunes in warm, late evening light with stormy evening sky, Bamburgh, Northumberland, UK. March 2014
Chlorophyte or green alga (Volvox aureus) in pond water. UK. The larger spheres are approximately 0.5mm in diameter. Digital composite image
Western red cedar tree (Thuja plicata) detail of bark, The Big Tree Trail, Meares Island, Clayoquot Sound, Vancouver Island, British Columbia, Canada
Herb robert (Geranium robertianum) flower, Coombe Valley, Cornwall, UK, May
Aurora Borealis (Northern Lights) over the Klondike River, Yukon Territories, Canada, September 2013
Beech (Fagus sylvatica) woodland in spring, at dawn, Peak District National Park, Cheshire, UK, May
Track through fir forest in autumn, Vosges mountain, France, October
Black Bear (Ursus americanus) sleeping in a Sitka Spruce tree (Picea sitchensis) alongside Anan Creek, Alaska, July
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Aerial view after sunrise, showing the sun reflecting off water that has flowed into Lake Eyre South and formed a pond; the water being the result of recent uncommonly high local desert rainfall
Goldfinch (Carduelis carduelis) feeding on teasel in falling snow, Leicestershire, UK, February
Bearded reedling (Panurus biarmicus) female feeding on reed seeds covered in frost, Espoo, Finland, January
Coastal agave (Agave shawii) leaves. Near Bahia de Los Angeles, Baja California, Mexico
Species rich alpine meadow with Orange lily (Lilium bulbiferum), Meadow clary (Salvia pratensis) and Yellow rattle (Rhinathus sp)
RF - White-legged damselfly (Platycnemis pennipes) peering over Oxeye daisy (Leucanthemum vulgare) flower. The Netherlands. August
Siberian tiger (Panthera tigris altaica) female territorial marking by rubbing cheek against ancient Manchurian fir tree, Land of the Leopard National Park, Far East Russia, November
Common oak (Quercus robur) fowers shedding pollen. Surrey, UK. April
Small Tortoiseshell Butterfly (Aglais urticae) feeding on scabious flowers in hay meadow, Wensum Valley, Norfolk, England, UK, July
Buckholt Wood is a beech (Fagus sylvatica) wood and part of the Cotwolds Commons and Beechwoods National Nature Reserve, on the scarp slope of the Cotswolds near Birdlip, Gloucestershire
Scottish primrose (Primula scotica) native only on the northern coast of Scotland and Orkney, here growing on grasslands at Dunnet Head, Caithness, England, UK, July
Early or English gentian (Gentianella anglica) a scarce plant, endemic to England, growing here at Durlston Head in Dorset, England, UK, May
Mother of pearl (Pleuroptya ruralis) moth nectaring on Ragwort (Jacobaea vulgaris) at night. Surrey, England, UK. July
Cedar (Cedrus sp), male catkins releasing pollen. Surrey England, UK
Garden bumblebee (Bombus hortorum) queen nectaring on Comfrey (Symphytum Hidcote Pink ) flowers. In garden, Surrey, England, UK. March
Rice (Oryza sativa) terraces. Jatiluwih Green Land, Bali, Indonesia. 2015
Palms growing in front of Rice (Oryza sativa) terrace. Jatiluwih Green Land, Bali, Indonesia. 2015
American lobster (Homarus americana) in eelgrass (Zostera marina). Nova Scotia, Canada. July
Red squirrel (Sciurus vulgaris) sitting on fallen tree in pine woodland, Glenfeshie, Cairngorms National Park, Scotland, UK, August