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Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Mallard (Anas platyrhynchos) shaking its wings with slow shutter speed to reveal movement, Martin Mere WWT Reserve, Lancashire, UK
Mute swan (Cygnus olor) cygnet swimming. Richmond Park, London, UK. May
Honey bee (Apis mellifera) approaching hive entrance, with trails of departing bees visible. Urban garden, Bristol, UK, May
Honey bee (Apis mellifera) flying close to hive entrance, with flight trails visible. Urban garden, Bristol, UK, May
RF - Alpacas (Vicugna pacos) group of three, Newton Farm, Inverarity, Angus, Scotland, UK, May. (This image may be licensed either as rights managed or royalty free.)
Kitten looking out of window, France
Borage (Borago officinalis) in flower with white background, Brechin, Scotland, UK, June
Mahogany treefrog (Tlalocohyla loquax) clasping onto leaf stem, view from below. Las Guacamayas Biological Station, Laguna del Tigre National Park, El Peten, Guatemala
Blue-green cyanobacteria (Trichodesmium thiebautii) Sargasso Sea, Bermuda
Compsite of orchid bees (Euglossa sp. ) photographed on black in cloud forestCompsite of orchid bees (Euglossa sp.) photographed on black in cloud forest, Choco region, Northwestern Ecuador
Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas
Portrait of a Ghost bat (Macroderma gigas) with dark background, captive, Adelaide Zoo, Australia
Bramble leaves (Rubus fruticosus) and bracken fronds changing colour in autumn
Kaleidoscopic pattern of Virginia creeper (Parthenocissus quinquefolia) in autumn on a white background
Kaleidoscopic image of Bramble leaves (Rubus fruticosus) and bracken fronds changing colour in autumn, on white background
Gyre flyingfish (Prognichthys glaphyrae) photographed in the Sargasso Sea, Atlantic Ocean
A larval stage slipper lobster (Scyllaridae) photographed in the Sargasso Sea, Atlantic ocean, International Waters
Californian sea lion (Zalophus californianus) swimming in Southern sea palm (Eisenia arborea) and Giant Kelp (Macrocystis pyrifera) forest, San Benitos Islands
Hibiscus (Hibiscus sp) flower with stigmas and pollen covered stamens. Focus stacked
Orange bush monkeyflower (Mimulus aurantiacus). Native to California and Oregon, USA. Focus stacked
Crocus (Crocus sp) flower from above, closing in evening. Focus stacked
Water tulip (Cypella aquatilis) from above, in garden pond. Native to Brazil
Daffodil (Narcissus sp) opening, view from above. Focus stacked
Peach leaved bellflower (Campanula persicifolia) dissection, petals removed to show how stamens transfer pollen to style via secondary pollen presentation in bud stage. Focus stacked
Evening primrose (Oenothera glazioviana) flowering at dusk in garden, Surrey, England, UK. August
Winter aconite (Eranthis hyemalis) flower and bud. Ring of nectaries formed from petals. In garden, Surrey, England, UK. February
Thorowax (Bupleurum rotundifolium) umbel with green bracts, backlit in studio
Elephant yam flower (Amorphophallus paeoniifolius) spadix surrounded by spathe. Cultivated in glasshouse
Masterwort (Astrantia sp) with pink bracts and tiny florets. Cultivated in garden, Surrey, England, UK. Controlled conditions
Spider flower (Cleome hassleriana) from above, flowers fading with age. Native to South America
Foxglove (Digitalis purpurea) flower, backlit with stamens and style visible through corolla. Dark spots on lower lip attract bumblebee pollinators. Pollinated by bumblebees with long proboscis
Love-in-a-mist (Nigella damascena) stamens fluroescing in UV light. Ring of nectaries are dark red with blue UV reflective hairs
Flowering rush (Butomus umbellatus) with stamens and stigma, nectar globules at base of carpels. Surrey, England, UK. Focus stacked
Orange-petaled passionflower (Passiflora aurantia) changing colour from cream to orange. Cultivated in glasshouse, Kew Gardens, London, England, UK. Native to New Guinea and northeast Australia
Afghanistan Juno iris (Iris cycloglossa) with yellow nectar guides. In cultivation, Kew Gardens, London, England, UK. Native to Afghanistan
Christmas rose (Helleborus niger) flower, pollen on dehisced anthers and on petals fluorescing in UV light. Nectar ring appearing red. Focus stacked
Blue passionflower (Passiflora caerulea). Flower is pollinated by larger bees. Focus stacked
Large-flowered evening primrose (Oenothera glazioviana). Bulls eye in centre of flower fluorescing in UV light guides pollinators to nectaries. At night in garden. Surrey, England, UK
Spider flower (Cleome sp) with long gonophores. View from above. Cultivated in garden, Surrey, England, UK. Native to South America
Silver Y moth (Autographa gamma) nectaring on Evening primrose (Oenothera glazioviana) with proboscis extended. At night in garden. Surrey, England, UK. August
Hazel (Corylus avellana) catkins dispersing pollen with female flower above on branch. Controlled condition.s Surrey, England, UK
Yellow mariposa lily (Calochortus luteus) with dark nectar guides. Cultivated in glasshouse, Surrey, England, UK. Endemic to California
Winter aconite (Eranthis hymenalis) in UV light, pollen on outer stamens that have dehisced fluorescing. Focus stacked
Tulip tree (Liriodendron tulipifera) flower with orange band to attract bees, produces copious nectar. In visible light. Surrey, England, UK. Native to USA. Focus stacked. Sequence 1/2
Handkerchief / Dove tree (Davidia involucrata) inflorescence, stamens with green ovary and multi--rayed style beneath. Native to Sichuan, northwest Yunnan, Guizhou and Hubei, China
Cremanthodium (Cremanthodium brunneopilosum). In wet meadow between Barkham and Ruo Er Gai, Sichuan Province, China
Shoal of three Bigfin reef squid (Sepioteuthis lessoniana) in open water at night. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea