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Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru
Trap-jaw ant (Odontomachus hastatus) with mandibles open, Los Amigos Biological Station, Peru
Hoatzins (Opisthocomus hoazin) perched in tropical rainforest, Tambopata Reserve, Peru, South America
Trap-jaw ant (Odontomachus chelifer) portrait, Wayqecha, Peru
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Red-and-Green Macaws (Ara chloropterus), feeding at the wall of a riverside clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Long-tailed potoo (Nyctibius aethereus) camouflaged on roosting perch in lowland rainforest, Manu Biosphere Reserve, Peru
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
Sphinx hawk moth (Hemeroplanes triptolemus) caterpillar, snake mimic species, Amarakaeri Communal Reserve, Maijuna, Rio Napo, Peru
Peruvian Tarantula (Pamphobeteus sp. ) adult, walking over Humming Frog (ChiasmocleisPeruvian Tarantula (Pamphobeteus sp.) adult, walking over Humming Frog (Chiasmocleis royi) without preying on it. Los Amigos Biological Station, Madre de Dios, Amazonia, Peru
Laguna 69, Huascaran National Park, Andes, Peru, November 2013
Giant waxy monkey frog (Phyllomedusa bicolor). Lowland Amazon rainforest, Manu Biosphere Reserve, Peru
River running through cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Dobsonfly female (Corydalinae) in cloud forest, Manu Biosphere Reserve, Peru
Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
White witch moth (Thysania agrippina) camouflaged against bark of tree. Manu Biosphere Reserve, Amazonia, Peru
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Red-and-green macaw (Ara chloropterus) flock feeding on wall of clay lick. Manu Biosphere Reserve, Amazonia, Peru
Hoatzin (Opisthocomus hoazin), pair on floating log in Cocha Salvador ox-bow lake. Manu Biosphere Reserve, Peru
RF - Maidens veil / Bridal veil fungus (Phallus indusiatus) with indusium fully formed, on rainforest floor. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Tufted / Brown capuchin (Cebus apella), female with baby on back, sitting in tree, mid-altitude montane forest, Manu Biosphere Reserve, Peru
Spectacled caiman (Caiman crocodilus) in shallow water, Amazon rainforest, Loreto, Peru
Blessed poison frog (Ranitomeya benedicta) male, carrying tadpoles on its back, transporting them to water, Yurimaguas, Alto Amazonas, Peru
Mealy Parrots (Amazona farinosa) feeding at the wall of a clay lick. Blanquillo Clay Lick, Manu Biosphere Reserve, Peru. November
Emperor tamarin (Saguinus imperator ) portrait, captive, occurs in Peru and Bolivia. Green foliage digitally added
Brown-mantled tamarin / Saddleback tamarin (Saguinus fuscicollis) climbing down tree trunk, Tambopata, Madre de Dios, Peru
Red-and-green macaw (Ara chloropterus) flock feeding at wall of clay lick, flying and perched in trees. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Collared peccary (Pecari tajacu) standing on the rainforest floor. Madre de Dios, Peru. March
Emperor Tamarin (Saguinus imperator) in a rainforest clearing. The animal is one of many being studied, hence the metal band hanging from its neck. Madre de Dios, Peru. March. Cropped
Golden-collared Tanager (Iridosornis jelskii), in the Peruvian cloud forest. Wayqecha Biological Research Station, Peru. April. Cropped
Purus red howler monkey (Alouatta puruensis) sitting in the rainforest canopy. Madre de Dios, Peru. April. Cropped
Dusky Titi monkey (Callicebus moloch) family troop sitting on a fallen tree in the Peruvian Amazon, with one grooming another. Madre de Dios, Peru. March
Hoatzin (Ophisthocomus hoazin) perched on wooden bar overlooking oxbow lake. Manu National Park, Madre de Dios, Peru. September
Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Paracas Beach, Paracas National Reserve, Ica Region, Peru. November 2013
Oreocallis grandiflora (Proteaceae family), Manu Cloud forest at 3500 metres altitude, Peru. November
Green-banded urania moth (Urania leilus) moths drinking salts from mineral-rich river clay, a behaviour know as puddling. On the banks of the Manu River, Manu Biosphere Reserve, Amazonia, Peru
Amazonian Horned Frog (Ceratophrys cornuta) camouflaged amongst leaf litter on lowland rainforest floor, waiting to ambush passing prey. Manu Biosphere Reserve, Amazonia, Peru. November
Three-striped poison frog (Ameerega trivittata) amongst leaf litter on lowland rainforest floor. Manu Biosphere Reserve, Amazonia, Peru. November
Nictitating membrane covering eye of Map Treefrog (Hypsiboas geographicus). The membrane affords camouflage and protection from dryness whilst allowing the frog to see
Andean Cock-of-the-rock male {Rupicola peruvianus} at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Brown / Tufted Capuchin (Cebus apella) family group in cloud forest, Manu Biosphere Reserve, Peru
Katydid / Bush Cricket (Tettigoniidae) camouflaged amongst cloud forest understory vegetation. 1600 metres altitude, Manu Biosphere Reserve, Peru. November
Crab Spider (Thomisidae) waiting in an orchid to ambush its prey. Cloud forest, Manu Biosphere Reserve, Peru. November
Bush cricket or katydid (Tettigoniidae) with spines, camouflaged amongst moss. Forest interior of mid-altitude montane rainforest, Manu Biosphere Reserve, Peru
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Aerial view of lowland Amazonia rainforest in morning mist. Manu Biosphere Reserve, Amazonia, Peru. November 2017
Bell glass frog (Cochranella nola) from below, photographed on a pane of glass in Manu Biosphere Reserve, Peru
Yellow crowned parrot (Amazona ochrocephala) at clay lick, Tambopata Research Centre, Peru
Detail of White Witch Moth (Thysania agrippina) in cloud forest, Manu Biosphere Reserve, Amazonia, Peru
Lupuna tree (Chorisia insignis) in lowland rainforest, Panguana Reserve, Huanuca province, Amazon basin, Peru
Reticulated tree frog (Dendropsophus reticulatus) portrait, Loreto, Peru
Imitating poison frog (Ranitomeya imitator) resting on leaf, Tarapoto, Peru. Cropped
Golden-tailed sapphire (Chrysuronia oenone) in flight. Pilcopata, Cuzco, Peru. May
Aguaje palm swamp (Mauritia flexuosa) Maijuna Indigenous Community, Rainforest, Sucusari, Rio Napo, Loreto, Peru. January 2013
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Red-and-green Macaws (Ara chloroptera) in rainforest, Tambopata National Reserve, Peru, South America
South American Sealion, (Otaria byronia) breeding colony. Punta San Juan Reserve, (Reserva Nacional de Islas, Islotes y Puntas Guaneras) Peru
Many-coloured rush tyrant (Tachuris rubrigastra) perched on reed stem at the waters edge. Lake Titicaca, Puno, Peru. September. Cropped
Peruvian Spider Monkeys (Ateles chamek) in the canopy of the Peruvian Amazon, at sleeping roost. Madre de Dios, Peru. April
Pink-toed tarantula (Avicularia avicularia), female waiting in ambush outside daytime lair in palm tree trunk. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Lava lizard (Microlophus peruvianus) juvenile peering out of eye of dead sea lion. Paracas National Reserve, Peru
Whip scorpion (Heterophrynus elephas) hunting for food on a large tree root of the rainforest. Los Amigos Biological Station, Peru
Black spider monkey (Ateles chamek) calling, captive, Peru
White lipped peccary (Tayassu pecari) on forest floor, Lago Preto Conservation Concession, Yavari River, Peru
Black spider monkey (Ateles chamek) portrait, captive, Peru
Aerial view of Amazon Rainforest, Peru, July 2015
Aerial view of Mouth of the Yavari-Mirin River entering Yavari River and Amazon Rainforest, Peru, July 2015
Aerial view of Amazon Rainforest and the Yavari River, and oxbow lake, Peru. July 2015
Scarlet macaw (Ara macao) in rainforest Tambopata National Reserve, Peru
Golden-mantled tamarin (Saguinus tripartitus) captive
Manu Road with small waterfall, Manu National Park, Madre de Dios Peru. August 2013. Road spans from Cuzco to the Madre de Dios River and Manu
Spanish cedar (Cedrela odorata) tree, Manu National Park, Peru
Fisheye view of Cocha Cashu Lake, Manu National Park, Peru
Portrait of an Emperor Tamarin (Saguinus imperator) mother with baby. Captive. Endemic to Peru
Margay (Leopardus wiedii) portrait, semi-free-ranging, Peruvian Amazon, Peru
Red Howler Monkey (Alouatta seniculus) hanging by tail with non-prehensile tailed Peruvian red uakari monkey (Cacajao calvus ucayalii)
Giant monkey frog / Blue and yellow frog (Phyllomedusa bicolor) on branch, Tambopata, Madre de Dios, Peru
Green jararaca / Two striped forest pit viper (Bothriopsis bilineata) on branch, Tambopata, Madre de Dios, Peru
Emerald tree boa (Corallus batesii) juvenile hanging on a branch. Tambopata, Madre de Dios, Peru
Hoatzin (Opisthocomus hoazin) chick perched on branch showing claws on wings. The claws are a vestigial feature which is lost by adulthood. Rio Yavari, Peru (non-ex)
Emerald tree Boa (Corallus batesii) close up portrait of juvenile whilst hanging on a branch. Tambopata, Madre de Dios, Peru
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
Jaguar (Panthera onca) portrait, Tambopata, Madre de Dios, Peru
