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Wandering albatross (Diomedea exulans) in flight over the ocean, Drake Passage, Southern Ocean
Atlantic puffin (Fratercula arctica) close up of head, Fair Isle, Shetland, Scotland, UK, July
Atlantic Puffins (Fratercula arctica) bill rubbing, Skomer Island, Wales, UK, April
Galapagos penguin (Spheniscus mendiculus) underwater, Bartholome Island, Galapagos Islands, East Pacific Ocean
White-vented storm petrel (Oceanites gracilis galapagoensis) running across water, offshore Cape Berkeley, Isabela island, Galapagos
Red-billed tropicbird (Phaethon aethereus) in flight, Plazas Island, Galapagos
Atlantic Puffin (Fratercula arctica) one landing among resting group, Sule Skerry, Scotland, UK, July
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
Gentoo penguins (Pygoscelis papua) group walking along snow, Cuverville Island. Antarctic Peninsula, Antarctica
Emperor penguin (Aptenodytes forsteri) tobogganing, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Northern gannet (Morus bassanus) in flight over colony, Quebec, Canada, August
Gannet (Sula bassana) diving, July UK Shetland
Gentoo Penguin (Pygoscelis papua) coming out of the sea, Cuverville Island, Antarctic Peninsula, Antarctica
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
Atlantic puffin (Fratercula arctica) amongst flowering Sea campion (Silene uniflora), Skomer Island, Pembrokeshire, Wales, UK. May
Emperor penguin (Aptenodytes forsteri) close up, Amanda Bay, Prydz Bay, Ingrid Christensen Coast, East Antarctica, November
Waved albatross (Phoebastria irrorata) head portrait, Espanola Island, Galapagos, South America
Laysan albatross (Phoebastria immutabilis) chick in the nest, Guadalupe Island Biosphere Reserve, off the coast of Baja California, Mexico, February
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Wandering albatross (Diomedea exulans) flying on the open ocean, Drake passage, Antarctic Peninsula, Antarctica
Short tailed albatross (Phoebastria albatrus) subadult with fish-hook and monofilament line embedded in throat. Tsubamezaki, Torishima Island, Japan. December
Sooty albatross (Phoebetria fusca) nesting amongst Blechnum palmiforme tree ferns. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display sequence. Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Erect-crested penguins (Eudyptes sclateri) pair. Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
White-vented storm petrel (Oceanites gracilis galapagoensis) walking on surface of water whilst foraging, Galapagos
Nazca booby (Sula granti) portrait, Galapagos, Ecuador
Great frigatebird (Phaethon aethereus) male at nest site, with gular sac inflated to attract females. Genovesa Island, Galapagos, May
Fulmar (Fulmarus glacialis) nesting amongst flowering thrift (Armeria maritima), Great Saltee Islands, County Wexford, Republic of Ireland. May
Emperor penguin (Aptenodytes forsteri), tobogganing, Snow Hill Island, Antarctic Peninsula
Atlantic puffic, (Fratercula arctica) portrait. Machias Seal Island, Bay of Fundy, New Brunswick, Canada, August
Atlantic yellow-nosed albatross (Thalassarche chlororhynchos) courtship display, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway
Westland petrel (Procellaria westlandica) in flight over sea, showing the upperwing. Kaikoura, South Island, New Zealand. April. Vulnerable species
Little blue / fairy penguin (Eudyptula minor) coming ashore to breeding area after dusk, Phillip Island Nature Park, Victoria, Australia
Tufted puffin (Fratercula cirrhata) in breeding plumage. St. Paul Island, Alaska. July
Horned puffin (Fratercula corniculata) close-up portrait, St. Paul Island, Pribilofs, Alaska, USA, July
Wilsons Storm Petrel (Oceanites oceanicus) feeding whilst walking on water Atlantic ocean, Madeira Portugal, August
Gannet (Morus bassanus) adult wing stretching and looking up, Shetland Islands, Scotland, UK, September
Atlantic Puffins (Fratercula arctica) three resting on a snow bank, Hornoya, Varanger, Norway, April
Atlantic puffin (Fratercula arctica) in flight, Isle of Lunga, Treshnish Isles, Scotland, June
Northern gannets (Morus bassanus) diving for fish. Bempton, Yorkshire, UK. August
Snow petrel (Pagodroma nivea) flying, Antarctica, November
Humboldt penguin (Spheniscus humboldti) group gathering at landing site. Tilgo Island, La Serena, Chile. Vulnerable species
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Yellow-eyed Penguin (Megadyptes antipodes) walking to nest hidden inland in dense vegetation. Sandy Bay, Enderby Island, Auckland Group, New Zealand Sub Antarctic Islands