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Cheetah (Acinonyx jubatus) running, Kalahari Desert, Botswana
Swallowtail (Papilio machaon) in flight, Finland, Europe
Cheetah (Acinonyx jubatus) mother and juveniles, Masai-Mara Game Reserve, Kenya. Vulnerable species
Red Sea anemonefish (Amphiprion bicinctus) in sea-anemone, Red Sea
Glass Frog (Cochranella mache) portrait, Ecuador, Endangered species
Sumatran tiger (Panthera tigris sumatrae) close-up head portrait, captive
Chicken wire coated in hoar frost. Peak District National Park, Derbyshire, UK, December 2012
Ten-spotted ladybird (Adalia decempunctata) on Common Water-plantain (Alsima plantago-aquatica) Belgium, June
Blackcap (Sylvia atricapilla) Uto Finland May
Humpback Whale (Megaptera novaeangliae) female. Tonga, South Pacific, September
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Bengal Tiger (Panthera tigris tigris) mother and cub, Pench National Park, Madhya Pradesh, India, taken on location for Tiger - Spy in the Jungle
Bengal Tiger (Panthera tigris) sub-adult, approximately 17-19 months old, lit by morning light, stalking. Endangered. Bandhavgarh National Park, India. Non-ex
Bengal Tigers (Panthera tigris tigris) resting in the shade national park, India, taken on location for Tiger - Spy in the Jungle March 2007
Playful tabby kitten, Stanley, 6 weeks
Amphipod (Iphimedia obesa) on top of a colony of Deadmans fingers (Alcyonium digitatum) coral, Loch Carron, Ross and Cromarty, Scotland, UK, April
Starlings (Sturnus vulgaris), adults perched on branch in winter feeding on apple. Lorraine France. 3rd Prize in the Melvita Nature Images Awards competition 2013
Female Jaguar (Panthera onca), captive, occurs in Southern and Central America
Leopard (Panthera pardus) resting in acacia tree, Samburu National Reserve, Kenya, Africa
Wood patterns in Lodgepole pine (Pinus contorta) Yellowstone National Park, Wyoming, USA, January
Antelope Canyon, a slot canyon with eroded sandstone patterns, Navajo Tribal Park, Arizona, USA, December 2012
Song thrush (Turdus philomelos clarkei) portrait, Hungary, May
Wild male Jaguar (Panthera onca palustris) stalking along the bank of the Cuiaba River in late afternoon sun light. Northern Pantanal, Brazil
Long-beaked Coralfish (Chelmon rostratus) eye, Inanuran Island, Danajon Bank, Central Visayas, Philippines, April
Rough Collie, bitch, 9 months in straw
Springbok (Antidorcas marsupialis) herd, Kgalagadi Transfrontier Park, South Africa
Whaleshark (Rhincodon typus) swimming and filtering fish eggs from the water. A behaviour known as ram-feeding. Isla Mujeres, Quintana Roo, Yucatan Peninsular, Mexico. Caribbean Sea
Portrait of a Spiny-chest Frog (Alsodes norae) endemic to Oncol Park, Chile, January
Male Long-tailed duck (Clangula hyemalis), Batsfjord, Norway, March
RF - Red fox (Vulpes vulpes) in dappled light on snow, Yellowstone National Park, USA. February
Leopard (Panthera pardus) back of head, Samburu National Reserve, Kenya
Leopard (Panthera pardus) portrait, Samburu National Reserve, Kenya
Leopard (Panthera pardus) female walking on fallen tree trunk, Samburu game reserve, Kenya
Apollo butterfly (Parnasius apollo) feeding on flower, Mount Terminillo, Rieti, Lazio, Italy, July 2011
Peruvian screech-owl (Megascops roboratus) half face close up portrait, Macara, Loja, Ecuador
Striped vinesnake (Philodryas argentea) Yasuni National Park, Orellana, Ecuador
Giraffe (Giraffa camelopardalis) close up of pattern Chobe River, North-West District, Botswana
Barred owl (Strix varia) stretching wings, Corkscrew Swamp Audubon Sanctuary, Florida, USA, March
Porcelain fungus (Oudemansiella mucida) gills. Peak District National Park, Derbyshire, UK. October
Badger (Meles meles) sub-adult badger among some ragwort, Derbyshire, England, UK, July. Highly commended in the Habitat category of the BWPA competition 2014
RF- Puffins (Fratercula) displaying. Skomer Island, Wales, UK. April
Red grouse (Lagopus lagopus scotica) sitting in flowering heather, Peak District, UK, September
Golden damselfish (Amblyglyphidodon aureus) hiding amongst the branches of red whip coral (Ctenocella sp.) Misool, Raja Ampat, West Papua, Indonesia
Cleaner wrasse (Labroides dimidiatus) swims out from the gill of a Blue spotted pufferfish (Arothron caeruleopunctatus). East of Eden, Similan Islands, Thailand. Andaman Sea, Indian Ocean
A portrait of a small Candy stripe shrimp (Lebbeus grandimanus) in front of its host pink anemone (out of focus because of the magnification)
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
Photo showing detail of the underside and mount of a honeycomb sea star (Pentaceraster alveolatus) with two tiny shrimps (Periclimenes soror). Bitung, North Sulawesi, Indonesia
RF - Bluestreak cleaner wrasse (Labroides dimidiatus) tending to Ribboned sweetlips (Plectorhinchus polytaenia). Yillet Kecil, Yillet Islands, Misool, Raja Ampat, West Papua, Indonesia