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Silhouette of Common peafowl (Pavo cristatus) perched in a tree at dawn. Keoladeo Ghana National Park, Bharatpur, Rajasthan, India
Black-and-white Border Collie bitch lying chin on floor with black-and-white kitten
European common frog (Rana temporaria) on rock by waterfall, near Lac des Gloriettes dam, Pyrenees National Park, France, June
Black veined white butterfly (Aporia crataegi) on grass, Gavarnie-Gedre, Pyrenees National Park, France, June
Great Blue shark (Prionace glauca) profile portrait viewed from slightly above, Pico Island, Azores, Portugal, Atlantic Ocean
Smooth coated Jack Russell Terrier, black, tan and white, female with two 8 week puppies
Banded demoiselle (Calopteryx splendens) group of four on plant stems, River Leijgraaf, Nijmegen, the Netherlands, August 2013
Kingfisher (Alcedo atthis) perched on branch, Black Forest, Baden-Wurttemberg, Germany
Female Snowy owl (Nyctea scandiaca) backlit surrounded by blowing snow, near Georgian Bay, Ontario, Canada
Great egret (Ardea alba) in snow, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. March
Adult Helmet Vanga (Euryceros prevostii) incubating a clutch of 3 eggs. Nest constructed in the fork of a Birds Nest Fern (Asplenium sp.) Masoala National Park, north east Madagascar
European Brown Bear (Ursus arctos arctos), a large male taking a rest in a cloud of mosquitoes. Kajaani, Finland. June
American Red Fox (Vulpes vulpes) cubs playing with its mother. Grand Teton National Park, USA, June
Blackbird (Turdus merula) male foraging on snow covered ground, during a blizzard. Derbyshire, UK. January (non-ex)
Chipping Sparrow (Spizella passerina) adult on ice covered Possum Haw Holly (Ilex decidua) berries. New Braunfels, San Antonio, Hill Country, Central Texas, USA, January
Grizzly bear (Ursus arctos) watching for Salmon at top of waterfall, Brooks Falls, Katmai National Park, Alaska, USA. July
Five Grizzly bears (Ursus arctos) standing at top of waterfall fishing for Salmon, Brooks Falls, Katmai National Park, Alaska, USA. July
Female Red fox (Vulpes vulpes) waiting to be fed at the back entrance of a house, Hungary. January
Stockdove (Columba oenas) shadow on the trunk of beech tree. Black Forest, Germany, April
Willow grouse (Lagopus lagopus) Kiilopaa, Inari, Finland, January
Long-jawed orbweaver (Metellina segmentata) in web at sunset. Dunwich Heath, Suffolk, UK, September
Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
European robin (Erithacus rubecula) adult with autumn leaves in background, Punkaharju, Etela-Savo / Southern Savonia, Ita-Suomi / Eastern Finland, Finland. September
Male impalas (Aepyceros melampus) standing in heavy rain, Masai-Mara Game Reserve, Kenya. February
Portrait of male Waterbuck (Kobus ellipsiprymnus) in the rain, Masai-Mara game reserve, Kenya
Four Yellow-billed storks (Mycteria ibis) standing in water, Lake Nakuru, Nakuru National Park, Kenya
Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean
Three Short-finned pilot whales (Globicephala macrorhynchus) close to the surface against the light, Costa Rica. Pacific ocean
Great reed warbler (Arcocephalus arundinaceus) chick begging for food, with bokeh affect from water, Pusztaszer, Hungary, June
Marbled white butterfly (Melanargia galathea) resting on reed, Devon, UK, July
Northern Hawk-Owl (Surnia ulula) seen through branches, southwest Finland, February
Burchells zebras (Equus quagga burchellii) standing side by side. Etosha NP, Namibia
Rufous elephant shrews (Elephantulus rufescens) Botswana
Mountain Hare (Lepus timidus) feeding. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, March
Puffin (Fratercula arctica) stretching wings, late evening light. Shetland Islands, Scotland, July
Aquatic Warbler (Acrocephalus paludicola). Zvanets fen mire, Belarus, Europe. Vulnerable Species
Mallard duck (Anas platyrhynchos) drake swimming through reflection of house, Bude canal, Cornwall, UK. April 2015
Hanuman langur / Northern Plains Grey Langur (Semnopithecus entellus) drinking from a water tap at Ranthambore fort. Ranthambore National Park, Rajasthan, India
Owlfly sulphur (Libelloides coccajus) on flower, near Gavarnie, Pyrenees National Park, France, June
Willow grouse (Lagopus lagopus) in winter plumage, Utsjoki, Finland, April
Leopard (Panthera pardus) back of head, Samburu National Reserve, Kenya
Helmet Vanga (Euryceros prevostii) on nest incubating eggs. Masoala Peninsula National Park, north east Madagascar. IUCN Vulnerable Species
Spotted scorpionfish (Scorpaena plumieri) lying motionless and camouflaged in soft corals. West Palm Beach, Gulf Stream, West Atlantic Ocean, Florida, USA
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Row of Starlings (Sturnus vulgaris) perched on wire. Marais Breton, Vendee, French Atlantic Coast, November
Dipper (Cinclus cinclus) perched on moss-covered waterfall, Peak District NP, Derbyshire, UK, May, Highly Commended, Habitat category, British Wildlife Phototgraphy Awards (BWPA) competition 2011
Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October. Non-ex
