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Pair of European grey wolves (Canis lupus) interacting, Tromso, Norway, captive, April
Whooper swan (Cygnus cygnus) in landscape of Landmannalaugar, Iceland, June
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Oilbird (Steatornis caripensis) nocturnal birds roosting and nesting in along walls of deep, narrow gorge in farmland, Tandayapa Valley, 1500m altitude, Ecuador
Two Turtle doves (Streptopelia turtur) perched on a rusting iron rail in a rain shower, Essex, England, UK, June
Red deer (Cervus elaphus) stag and hind at dusk, Cheshire, UK September
European rabbit (Oryctolagus cuniculus) two young rabbits, or kittens, touch noses, Norfolk, UK, June. Non-ex
Rhinoceros hornbills (Buceros rhinoceros) pair flying over the canopy with mist hanging over lowland Dipterocarp rainforest in heart of Danum Valley, Sabah, Borneo (digitally modified image)
Noddy Turns (Anous stolidus) pair on Heron Island, southern Great Barrier Reef, Queensland, Australia
Two Canada geese (Branta canadensis) on a misty lake at dawn, UK
A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)
A pair of Australian Sea Lions (Neophoca cinerea) socialising in the shallows. Kangaroo Island, South Australia, Australia. Southern Ocean. November
Pair of European bee-eaters (Merops apiaster) with courtship offering of insect prey, Pusztaszer, Kiskunsagi National Park, Hungary
View of hikers from behind the Pericnik Waterfall, Slovenia, October 2018
Pair of Gannets (Morus bassanus) mutual preening, Bass Rock, Firth of Forth, Scotland, UK, June
Jamess flamingos {Phoenicoparrus jamesi} Laguna Colorado, Bolivia
Harvest mice (Micromys minutus), captive, UK, June
Northern gannets (Morus / Sula bassanus) portrait of individual with a courting pair in foreground, UK, Bass Rock, Scotland
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
Two Oleander Hawkmoths (Daphnis nerii) at rest on leaves
