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Nazca booby (Sula granti), pair billing on coastal cliff in evening light. Wolf Island, Galapagos. August 2016
Blue-footed booby (Sula nebouxii), group fishing underwater. Itabaca Channel, Santa Cruz Island, Galapagos
Aerial view of small coral island, Western Provinces, Solomon Islands
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Soft corals (Dendronephthya sp) in coral reef, West Papua, New Guinea
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Orange anemonefish (Amphiprion sandaracinos) in its host Mertens carpet sea anemone (Stichodactyla mertensii). Mabul Island, Sabah, Borneo, Malaysia. Celebes Sea / Sulawesi Sea
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Patagonian redfish (Sebastes oculatus) about 4cm, Comau Fjord, Patagonia, Chile
Mandarinfish (Synchiropus splendidus) Yap, Micronesia
Plankton at night, Palau, Philippine Sea
Marine iguana (Amblyrhynchus cristatus) in the surf, Black Beach, Floreana Island, Galapagos
Marine iguana (Amblyrhynchus cristatus) in water, Puerto Egas, Santiago Island, Galapagos
Galapagos lava lizard (Microlophus albemarlensis) sitting on Marine iguana (Amblyrhynchus cristatus) Punta Espinosa, Fernandina Island, Galapgos
Galapagos green turtle (Chelonia mydas agassizii) swims over a coral reef. Darwin Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Oceanic manta (Mobula birostris). Cousins Rock, San Salvador Island, Santiago Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Galapagos barnacle blenny (Acanthemblemaria castroi) looking out from its home in an old barnacle shell. Cape Marshall, Isabela Island, Galapagos National Park, Galapagos Islands. East Pacific Ocean
Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii
Clarks anemonefish (Amphiprion clarkii), pair tending to egg mass placed beside the protection of an anemone. Yap, Micronesia
RF - Polyps on gorgonian fan coral. West Papua, Indonesia
Variegated feather duster (Bispira variegata) Sabellidae. Puerto Galera, Philippines
RF - Orange skunk clownfish (Amphiprion sandaracinos) in anemone, Puerto Galera, Philippines
Magnificent feather duster worm (Protula magnifica) Puerto Galera, Philippines
Polyps on gorgonian fan coral. West Papua, Indonesia
Kaleidoscopic image of coral reef with Red sea whips (Ellisella ceratophyta). Misool, Raja Ampat, West Papua, Indonesia
Kaleidoscopic image of gorgonian coral. Misool, Raja Ampat, West Papua, Indonesia
Kaleidoscopic image of coral reef with soft corals (Dendropnethya) shoal of Yellow back fusilliers (Caesio teres) West Papua, Indonesia
Crinoid (Crinoidea) on artificial reef. Mabul, Malaysia
Acoel flatworms (Waminoa sp. ) on coral. Lembeh Strait, North Sulawesi, IndonesiaAcoel flatworms (Waminoa sp.) on coral. Lembeh Strait, North Sulawesi, Indonesia
RF - Coral reef with Soft corals (Dendronephthya sp) and fish, West Papua, Indonesia
Emperor shrimp (Periclemenes imperator) on a nudibranch (Ceratosoma sp. ) Lembeh StraitEmperor shrimp (Periclemenes imperator) on a nudibranch (Ceratosoma sp.) Lembeh Strait, North Sulawesi, Indonesia
Tube anemone (Cerianthus sp) Lembeh, Sulawesi, Indonesia
Gorgonian sea fan (Melithaea sp. ) with Lyretail anthias (Pseudanthias sqamipinnis)Gorgonian sea fan (Melithaea sp.) with Lyretail anthias (Pseudanthias sqamipinnis). Similan Islands, Andaman Sea, Thailand
Kaleidoscopic image of Bubble tip anemone (Entacmaea quadricolor). Raja Ampat, West Papua, Indonesia
Kaleidoscopic image of Feather stars (Crinoidea) and Gorgonian wrapper anemone.(Nemanthus annamensis). Indonesia. Western Pacific
A Kamchatka Brown Bear (Ursus arctos beringianus) standing in breaking waves. Pacific coast of Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Bottlenosed dolphin (Tursiops truncatus) porpoising during annual sardine run, Port St Johns, South Africa. June
RF - Shoal of Slender pinjalo snappers (Pinjalo lewisi) swimming over soft corals on coral reef. West Papua, Indonesia
Ring-tailed Cardinalfish (Apogon aureus) West Papua, Indonesia
Goldring tunicate (Clavelina robusta) Puerto Galera, Philippines
Gold-spotted Chromodorid nudibranch (Chromodoris aureopurpurea, Ninepin Group or Kwo Chau Islands)Gold-spotted Chromodorid nudibranch (Chromodoris aureopurpurea, Ninepin Group or Kwo Chau Islands, Sai Kung District of Hong Kong, China