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Trumpeter Swans (Cygnus buccinator) in winter morning mist, showing aggression during courtship behaviour, Mississippi River, Minnesota, USA, February
Quiraing, Trotternish Ridge, Isle of Skye, Scotland, December
Bluebells (Hyacinthoides non-scripta) flowering in oakwood, Perthshire, Scotland, UK, May
Flock of Sanderling (Calidris alba) flying over a rough sea, Berwickshire Coast, Scotland, UK, March
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
Red howler monkey (Alouatta seniculus) howling, captive
Ice cliffs of the Ross ice shelf, Antarctica
Black browed albatross flying over iceberg {Thalassarche melanophrys} South Georgia
Galapagos green turtle (Chelonia agassizii) surfaces to breathe. Floreana Island, Galapagos Islands, Ecuador. East Pacific Ocean
Wild gladiolus (Gladiolus illyricus) flowers, Menorca, Balearic Islands, Spain, Europe
Ancient dead Camelthorn tree (Vachellia erioloba) trees with red dunes and mist, Namib desert, Deadvlei, Sossusvlei, Namibia. August 2015
Boabab trees {Adansonia grandidieri} silhouetted at sunset. Morondava, Madagascar, October 2009
Snares-crested penguin (Eudyptes robustus) walking along rocks, Snares Island, New Zealand Sub-Antarctic Islands. Endemic and vulnerable species
Little blue / fairy penguin (Eudyptula minor) walking on pathway to nesting burrows, Neck Game Reserve, Bruny Island, Tasmania, Australia, December
Gentoo penguins (Pygoscelis papua) calling, Cuverville Island. Antarctic Peninsula, Antarctica
Gentoo Penguin (Pygoscelis papua) jumping out of the sea, Cuverville Island, Antarctic Peninsula, Antarctica
Gentoo Penguin (Pygoscelis papua) Wiencke Island, Antarctic Peninsula, Antarctica
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
Kimmeridge Bay ledges at sunset, Dorset, UK. September 2014
Tail fluke of a diving Humpback whale (Megaptera novaeangliae) misty coast in background, Alaska, USA. Gulf of Alaska, Pacific ocean
Colmers Hill in morning mist, near Bridport, Dorset, UK. September 2012
Avenue of Oak trees (Quercus) Eickelberg, Warnowtal, Germany, September
Blue sheep (Pseudois nayaur) Hemis National Park, Jammu and Kashmir, India
Mountain landscape, Svalbard, Norway, July
Ibex (Capra Ibex) fighting, Aiguilles Rouges National Nature Reserve, Haute-Savoie, France, June
Mountain Peaks at dusk, from left to right, Aiguille de Blaitiere, Aiguille de Cigeaux, and Aiguille du Fou. Chamonix, France, September 2012
Pacific hatchetfish (Argyropelecus affinis) portrait
RF - Harts tongue fern (Phyllitis scolopendrium) leaf unfurling and creating the shape of a heart, Cornwall, UK, May
RF- Gannets (Morus) displaying, Great Saltee, County Wexford, Ireland. May
Koala (Phascolarctos cinereus) mother with joey aged eight months, Queensland, Australia, captive
African Lion (Panthera leo) cub reaches for a moment of intimacy with its father, Northern Tuli Game Reserve, Botswana
Two newly fledged burrowing owl chicks (Athene cunicularia) one being groomed by its mother (far left) Pantanal, Brazil. WINNER
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Feral domestic rabbit (Oryctolagus cuniculus) juvenile running with dead leaf in mouth, Okunojima Island, also known as Rabbit Island, Hiroshima, Japan
Raccoon (Procyon lotor) standing up investigating camera, portrait, Stanley park, Vancouver, British Columbia, Cananda, September
Basset hound, portrait, resting head on back of car, eyes half closed, UK
Polar bear (Ursus maritimus) young male playing / rolling around in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Head portrait of Highland cow, Scotland, with tiny frond of bracken at corner of mouth, UK
Three Ring-tailed Lemurs (Lemur catta) sun basking at dawn. Berenty Private Reserve, southern Madagascar. (digitally stitched image)
RF - Close up of Greater stitchwort (Stellaria holostea) flower, Cornwall, UK. April
RF- Grape hyacinths (Muscari) and Daffodils (Narcissus sp) in flower, Norfolk, UK, March
Cornfield annual summer wildflowers growing on one of the plant charity Landlife sCornfield annual summer wildflowers growing on one of the plant charity Landlifes National Wildflower Farms, St Helens, Merseyside, UK, June
Giant Water Lily (Victoria amazonica) flower in a lagoon at Porto Jofre, off the Cuiaba River, northern Pantanal, Mato Grosso State, Brazil, South America
RF- Grey seal (Halichoerus grypus) pups on Norfolk Beach, England, UK, December
Japanese macaque (Macaca fuscata fuscata) mother with rare white furred baby, Jigokudani Valley, Nagano Prefecture, Japan. June
Baby ground squirrel (Xerus inauris) using its tail for shade, Kgalagadi Transfrontier Park, South Africa. Non-ex
RF- African elephant family (Loxodonta africana) crossing desert landscape. Namibia, Kaokoland. Endangered species
